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1.
Giant azhdarchids are perhaps the most spectacular pterosaurs known and, although they lack the elaborate headcrests or striking dentition that adorn other charismatic pterosaurs, the sheer enormity of their proportions has drawn a wealth of popular and scientific interest. Their remains, however, are so fragmentary that only limited interpretations of their anatomy and taxonomy can be made with certainty. Here, we report on recent investigations into giant azhdarchid taxonomy and size estimates that suggest both may be over-inflated.  相似文献   

2.
The Late Cretaceous (Santonian) continental vertebrate locality of Iharkút, western Hungary has provided numerous azhdarchid pterosaur remains including the recently described, Bakonydraco galaczi. Since the first report of these fossils, additional remains have been discovered that improve considerably our knowledge of some aspects of the anatomy of azhdarchid pterosaurs. New cranial material described here indicates an edentulous, non-crested premaxillary rostrum in Bakonydraco similar to that of Quetzalcoatlus and reveals that this rostrum was considerably thinner and more lightly built than the relatively massive, pointed mandibular symphysis. In addition, the contact surface of the upper and lower jaws of Bakonydraco at least in the symphyseal region was more similar to Tapejara wellnhoferi than to other azdarchids: the contact is irregular and the premaxilla does not fit closely the rostral part of the mandible. Among the postcranial material the atlas-axis complex possessing lateral pneumatic foramina is of great importance because it further supports the notion of interspecific variability of vertebral pneumaticity among pterosaurs.  相似文献   

3.
Fragmentary remains, including cervical vertebrae and limb bones, of a large pterosaur from the Upper Cretaceous of Tous, province of Valencia (Spain), are described. The material was recovered from lacustrine beds in the upper part of the Calizas y Margas de Sierra Perenchiza Formation, which is probably Maastrichtian in age. Six fragments of vertebrae allow a reconstruction of the anatomy of the mid-series cervicals of the animal. The general morphology of the cervical vertebrae is closely similar to that of the long-necked Azhdarchidae. Compared to other azhdarchids, the Valencia pterosaur shows minor differences from the genera Azhdarcho and Quetzalcoatlus, and is here provisionally referred to as Azhdarchidae indet. A wingspan of about 5.5 m is calculated by comparison with other known azhdarchids. This is the second azhdarchid pterosaur described from the Iberian peninsula. It confirms the wide distribution of this group of large pterosaurs at the end of the Cretaceous.  相似文献   

4.
Azhdarchid pterosaurs include the largest known flying animals, with the largest species reaching a potential mass of over 250 kg. Prior work suggests that several features of azhdarchid anatomy could be associated with a soaring-dominated lifestyle, including large size, burst-flapping adapted pectoral girdle and proximal forelimb, moderate to high wing aspect ratio, and exceptional pneumaticity.  相似文献   

5.
The partially articulated, distal portion of a left wing finger of a pterosaur from the Crato Formation of northeast Brazil has a T-shaped cross-section to the second and third phalanges. This cross-sectional shape is one of several characters diagnostic of the pterodactyloid pterosaur family Azhdarchidae (Unwin & Lu 1997). Until now, this family of pterosaurs was known exclusively from the Late Cretaceous. The specimen described here may be the first recorded azhdarchid from the southern hemisphere and the earliest recorded member of the family.  相似文献   

6.
Pterosaurs exhibit a large size range with the smallest approximately the size of a sparrow and the largest about the size of a small airplane. Based on studies of extant flying vertebrates and principles of allometry (Norberg & Rayner, 1987; Rayner, 1988), a reasonable assumption is that pterosaurs of different sizes used their wings differently during flight.  相似文献   

7.
A new specimen of pterosaurian metacarpal IV from the Early Cretaceous (Berriasian) Durlston Formation of Dorset, southern England, closely resembles those of the Chinese dsungaripterid Dsungaripterus weii Young, 1964 and the Central Asian azhdarchid Azhdarcho lancicollis Nessov, 1984. However, the new specimen lacks the thickened bony wall typical of Dsungaripteridae and is therefore regarded as azhdarchoid. An mcIV with a markedly offset distal condyle with well-developed diaphyseal constriction behind the distal condyle and asymmetrical condylar margins may be an autapomorphy of Dsungaripteridae + Azhdarchoidea, while the same condition with bone thickening may typify Dsungaripteridae. The new specimen also provides osteological evidence supporting claims for large wing-spanned pterosaurs in the Berriasian of southern England based previously only on ichnological finds. The new specimen suggests a wingspan of between 2.9 and 3.7 m and represents the largest pterosaur from the Berriasian.  相似文献   

8.
根据发现于辽西朝阳九佛堂组一具有部分头骨的不完整骨架,命名一新的帆翼龙--赵氏龙城翼龙(新属新种).头骨和牙齿形态显示龙城翼龙与宽齿帆翼龙的关系要比与其他翼龙的关系密切,龙城翼龙是目前辽西九佛堂组中发现的唯一的帆翼龙类.帆翼龙科原仅一属种,发现于英国早白垩世地层中,辽西早白垩世新帆翼龙的发现,不但扩大了其地理分布,也证明辽西是晚侏罗世至早白垩世翼龙辐射、分异的重要地区.这对研究翼龙的起源和演化有重要意义.  相似文献   

9.
白垩纪四足动物足印的生物地层学、生物年代学与遗迹相   总被引:3,自引:2,他引:1  
从全球范围来看,白垩纪四足动物的足印多数是非鸟恐龙与鸟类留下的痕迹;少量足印来自翼龙、鳄鱼、龟、哺乳动物和其他四足动物。白垩纪的足迹化石以东亚(尤其是中国和朝鲜)和北美西部的最为人所知。南美(主要是阿根廷和巴西)也有一定数量广泛分布的足迹化石,欧洲、非洲与澳大利亚的白垩纪足迹组合则鲜为人知。以白垩纪四足动物的足印记录为基础,我们对两个全球足印生物年代重新进行了检查。早白垩世生物年代以蜥脚类与鸟脚类的足迹为特征。晚白垩世生物年代中的蜥脚类足迹较少,但是鸭嘴龙、暴龙和角龙的足迹增多了。另外,白垩纪足印化石的记录中记载了许多重要的生物地层学信息,如北美白垩纪中期蜥脚类恐龙的消失,以及白垩纪末恐龙的绝灭。越来越多来自东亚的白垩纪足印记录使我们对更精细的地方性白垩纪足印生物年代学有了初步印象。因此,以地方性四足恐龙(包括鸟类)遗迹属的地层分布为基础,可以识别出三个或四个足印生物年代。种类丰富并具有地方性特色的东亚的白垩纪鸟类动物的遗迹群,可能指示白垩纪时东亚存在着一个独特而繁盛的鸟类动物群。以足印化石为基础的这一假说有待进一步的验证。  相似文献   

10.
The latest Cretaceous continental vertebrate faunas of the wider Transylvanian area figured prominently in discussions concerning the Cretaceous–Paleogene Boundary (K-Pg) events when they were first described by Nopcsa between 1897 and 1929, because they were assumed to be late Maastrichtian in age. Subsequently their age was reconsidered as early Maastrichtian, and were thus regarded of lesser importance in understanding the K-Pg boundary events in Europe and worldwide. Moreover, Transylvanian continental vertebrate assemblages (the so-called ‘Haţeg Island’ faunas) were often lumped together as a temporally restricted assemblage with a homogenous taxonomic composition. Recent fossil discoveries and more precise dating techniques have considerably expanded knowledge of the Transylvanian vertebrate assemblages, their ages, and their evolution. A synthesis of the available stratigraphic data allows development of the first comprehensive chronostratigraphic framework of the latest Cretaceous Transylvanian vertebrates. According to these new data, expansion of continental habitats and emergence of their vertebrate faunas started locally during the latter part of the late Campanian, and these faunas continued up to the second half of the Maastrichtian. During this time, long-term faunal stasis appears to have characterized the Transylvanian vertebrate assemblages, which is different from the striking turnovers recorded in western Europe during the same time interval. This suggests that there was no single ‘Europe-wide’ pattern of latest Cretaceous continental vertebrate evolution. Together, the available data shows that dinosaurs and other vertebrates were relatively abundant and diverse until at least ca. 1 million years before the K-Pg boundary, and is therefore consistent with the hypothesis of a sudden extinction, although this must be tested with future discoveries and better age constraints and correlations.  相似文献   

11.
Knowledge of the turtle fauna from the Lower Cretaceous of the Iberian Peninsula has been very limited until now. There are several fossil sites where Lower Cretaceous associations of continental vertebrates have been found. Although turtles have been identified in some of them, most of these specimens have not been studied, so the diversity is unknown. Among all these findings, the turtles from the Cameros Basin are considered particularly relevant, both in their abundance and diversity. Their study has allowed the identification of several taxa. At least one representative of Solemydidae and three taxa of Eucryptodira are recognized. This study establishes kinship and biogeographic relationships between the taxa in Cameros with those found in other Spanish fossil sites and with those of other European regions.  相似文献   

12.
季强  袁崇喜 《地质论评》2002,48(2):221-224
本文实事求是地介绍了道虎沟生物群的组成及相关地层的岩性,并根据长尾型嘴口龙类(Rhamphorhynchoidea)翼龙与短尾型翼手龙类(Pterodactyloidea)翼龙同期共生的特点,确信道虎沟生物群的地质时代为晚侏罗世。此外,本文还研究了道虎沟生物群中翼龙身上发育的皮肤衍生物,初步认为它们是原始羽毛(protofeathers),至少是与羽毛同源的(feather-homologous),这对于了解羽毛(广义)的演化历史和早期的形态结构将具有十分重要的意义。  相似文献   

13.
Although the European dinosaur succession during the latest Cretaceous and its relationship with the Cretaceous-Paleogene (K-Pg) mass extinction has been the focus of recent work, other continental vertebrates, such as crocodylomorphs, have received less attention. The European continental record of crocodylomorphs in general, and of eusuchians in particular, is relatively dense through the Maastrichtian until the K-Pg boundary. Traditionally it has been argued that continental crocodylomorphs were minimally impacted by the K-Pg extinction, but they were substantially affected in Europe with the disappearance of endemic eusuchians such as Hylaeochampsidae, Allodaposuchus and their close relatives, and non-eusuchians such as Doratodon or Theriosuchus. Despite extensive sampling in Danian continental deposits, only scarce fragmentary crocodylomorph remains have been cited. It is not until the late Paleocene and Eocene that a recovery in continental crocodylomorphs is observed. The presence of taxa such as planocraniids, the alligatoroids Diplocynodon and Hassiacosuchus, and stem crocodyloids during this period provide the first reliable continental records of Crocodylia in Europe and is best explained by post-extinction immigration from Asia or North America. By contrast, marine forms such as Thoracosaurus are found on both sides of the K-Pg boundary in Europe. The adaptation of these marine animals to different environments, from shallow seas to more transitional or fluvial environments, could be the key to their success and survival across the K-Pg boundary, as seen in other marine crocodylomorph clades such as Dyrosauridae.  相似文献   

14.
In the present paper, the fossil record of the archosaurs (dinosaurs, crocodylomorphs and pterosaurs) of the southern Pyrenees before the Cretaceous–Palaeogene (K–Pg) transition is revised. On the basis of this fossil record, a well-dated succession of dinosaurs and other archosaurs is established within polarity magnetochrons C30 and C29r. Almost 150 sites with dinosaur remains have been identified, containing hadrosauroid ornithopods, titanosaur sauropods and theropods, as well as egg sites and tracks. Fossil remains of dinosaurs and other archosaurs are abundant in C29r, disappearing abruptly near the top of the “Lower Red Garumnian” unit of the Tremp Formation. Thus this should be located very close to, or coinciding with the K–Pg boundary. These data suggest that the disappearance of the dinosaurs and other archosaurs was geologically abrupt in the southern Pyrenees, but to date there is no incontrovertible evidence of the presence of the impact level that marks the Cretaceous–Palaeogene boundary. Interestingly, what is highlighted in the southern Pyrenees is that the vertebrate-rich upper Maastrichtian continental sites were replaced by similar sedimentological facies characterized by the virtual absence not only of dinosaurs but also of any vertebrate remain throughout the lower Palaeocene. This could mean that the Danian terrestrial ecosystems of the southern Pyrenees took longer than other areas of the world to recover their biodiversity after the K−Pg extinction event.  相似文献   

15.
Some of the best records of continental vertebrates from the Cretaceous of Europe come from Romania, particularly two well-known occurrences of dwarfed and morphologically aberrant dinosaurs and other taxa that lived on islands (the Cornet and Haţeg Island faunas). Substantially less is known about those vertebrates living in the more stable, cratonic regions of Romania (and Eastern Europe as a whole), particularly during the earliest Cretaceous. We describe one of the few early Early Cretaceous fossils that have ever been found from these regions, the tooth of a large theropod dinosaur from Southern Dobrogea, which was discovered over a century ago but whose age and identification have been controversial. We identify the specimen as coming from the Valanginian stage of the Early Cretaceous, an incredibly poorly sampled interval in global dinosaur evolution, and as belonging to Carcharodontosauridae, a clade of derived, large-bodied apex predators whose earliest Cretaceous history is poorly known. Quantitative analyses demonstrate that the Romanian tooth shows affinities with a derived carcharodontosaurid subgroup, the Carcharodontosaurinae, which until now has been known solely from Gondwana. Our results suggest that this subgroup of colossal predators did not evolve vicariantly as Laurasia split from Gondwana, but originated earlier, perhaps in Europe. The carcharodontosaurine diversification may have been tied to a north-to-south trans-Tethyan dispersal that took place sometime between the Valanginian and the Aptian, illustrating the importance of palaeogeographic ties between these two realms during the largely mysterious early–mid Early Cretaceous.  相似文献   

16.
New fossils of dinosaurs and pterosaurs have been found in the conglomeratic facies of the Maungataniwha Member of the Tahora Formation (Campanian) of New Zealand. These derive from a large theropod, a sauropod, an ankylosaur and a pterosaur. Together with previously described material they indicate at least five, maybe seven, taxa from the Late Cretaceous terrestrial fauna of New Zealand. At that time New Zealand was an island near Antarctica, so this represents an insular, polar fauna.We assume a vicariance model for the origin of this fauna, which probably samples that present in Antarctica at the time New Zealand rifted away from it. The fauna differs from other well known Cretaceous Gondwana faunas in including an ankylosaur, but is similar to that from the Late Early Cretaceous of Queensland, Australia. The inclusion of both an ankylosaur and sauropod lend a relict, Early Cretaceous aspect to the fauna. There seems to be no particular similarity to known polar faunas from the Early Cretaceous of Victoria, Australia, or the Late Cretaceous of Alaska and Antarctica.Dinosaurs, both large and small, were capable of surviving in the cool to cold-temperate, seasonal climate of New Zealand at this time.  相似文献   

17.
近年来,在中国辽宁西部、内蒙古东南部、河北北部中侏罗世髫髻山组和九龙山组中发现了丰富的脊椎动物、无脊椎动物和植物化石,这一化石群被称为燕辽生物群。燕辽生物群已正式命名的脊椎动物26属28种,包括鱼类1属1种、两栖类4属4种、有鳞类1属1种、翼龙类10属12种、兽脚类恐龙5属5种、哺乳类5属5种。燕辽生物群的脊椎动物化石对研究长羽毛兽脚类恐龙的演化、翼龙类的分类演化、滑体两栖类的分异等具有重要意义。虽然辽宁西部及邻区燕辽生物群脊椎动物的多样性不及同一地区早白垩世的热河生物群,但它为认识这一地区晚中生代生物群的演替提供了极为重要的依据。  相似文献   

18.
Titanosaurs were a globally distributed group of sauropod dinosaurs. They had diverse forms and a wide-gauge stance, with a few of their species reaching immense sizes, such as Argentinosaurus huinculensis and Patagotitan mayorum (reaching >35 m in length). There are about 100 valid titanosaur species known so far, but most of the originally described species are no longer valid, due to the incomplete nature of fossil materials. Our understanding of titanosaur skull morphology is based on very few incomplete fragmented cranial materials and findings of the complete skull are even rarer. Understanding the skull morphology of extinct animals helps palaeontologists make deductions of feeding mechanisms and also provide an idea about their appearance when they were alive. Diversity in titanosaur skull morphology is greater than that of any other sauropod clade, indicating diversity in feeding mechanism among these dinosaurs. Titanosaurs were the last surviving clade of sauropod dinosaurs, occupying nearly every ecological niche around the world during the Late Cretaceous, and resulting in a rich diversity in this group. This article highlights diversity in the basic structure of sauropods with special emphasis on titanosaur skull morphology.  相似文献   

19.
Based on a nearly complete lower jaw from the Early Cretaceous of Liaoning Province, a new ctenochasmatid pterosaur: Liaoxipterus brachyognathus gen, et sp. nov. is erected. Liaoxipterus brachyognathus is different from any known pterosaurs with skulls preserved from western Liaoning Province and its peripheral areas in that the anterior part of the mandibular symphysis is expanded, being widest between the fourth alveolus of each side. Liaoxipterus brachyognathus is assigned to Ctenochasmatidae based on the following characters: the rounded anterior end of the lower jaw is spatulated and dorsoventrally flattened and marked heterodonty in the dentition is absent. It differs from other ctenochasmatid pterosaurs in having relatively small number of teeth. Liaoxipterus is distinguished from some ornithocheirids, which have expanded anterior parts of the mandibular symphyses, such as Anhanguera piscator,Coloborhynchus robustus in which the teeth of the new pterosaur are not as variable.  相似文献   

20.
Phylogenetic analyses of early pterosaur species are relatively new and contradictory. However, they imply a similar evolutionary history for early pterosaurs: a large divergence of all major early groups of pterosaurs before the first named pterosaur species appeared in the fossil record. This large, unsampled divergence may be a record of the initial adaptive radiation of the pterosaurs, but a recent comprehensive phylogenetic analysis reconstructs named Triassic pterosaur species grouped together in a monophyletic clade with all other major groups diverging later. Ten patterns that characterize adaptive radiations put forward by Gavrilets and Losos (2009) are assessed for congruence with the comprehensive phylogeny and two previous phylogenies to determine which are consistent with our notions of an adaptive radiation. Of the eight patterns that can be applied to pterosaurs, only six vary between pterosaur phylogenies. Of these six patterns, all apply to the three phylogenies with varying amounts of support. For many patterns, however, the comprehensive phylogeny has a greater magnitude, shorter time span, or is the only topology within which they can be assessed. Pterosaurs are characterized by an adaptive radiation no matter which phylogeny is considered, but congruence with the fossil record increases the support and information content of a phylogeny by reducing the time and area within which to reconstruct evolutionary history.  相似文献   

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