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1.
The orthophragminids in lower Bartonian Reineche Limestone member, a fossiliferous shallow-marine unit exposed in Cap Bon peninsula in Tunisia, are represented by 17 species assigned to Discocyclinidae Galloway 1928 and Orbitoclypeidae Brönnimann 1946. These taxa, associated with nummulitids and alveolinids, belong to the lineages of Discocyclina Gümbel 1870, Nemkovella, 1987, Orbitoclypeus Silvestri 1907, and Asterocyclina Gümbel 1870, described for the first time from north Africa lying at the southern margin of Tethyan ocean during Paleogene. We identified Nemkovella evae, previously not recorded in upper Lutetian/lower Bartonian and younger Eocene deposits of northern Tethyan platforms, and erected a new subspecies, N. evae reinechensis n. ssp. A comparison of Reineche orthophraminids, assigned to orthophragmines zone (OZ) 12 and shallow benthic zone (SBZ 17), to the well-described coeval assemblages at northern Tethyan platforms in Italy, Hungary, Turkey, and to those in Kutch Basin in the Indian subcontinent suggests that some species are confined to certain paleogeographic domains. Orbitoclypeus haynesi, the only orbitoclypeid and the most abundant orthophragminid in lower Bartonian deposits in Kutch, appears to be the most common orbitoclypeid in Reineche Limestone. In Europe, this species is not known and is replaced by Orbitoclypeus varians, the most common orbitoclypeid in middle Eocene of central Europe. Both species occur in varying proportions in marine successions in Turkey. Asterocyclina sireli, identified so far only in Turkey, occurs in Reineche Limestone and in lower Bartonian deposits in Kutch. This species is recorded for the first time in the Indian subcontinent. Relying on present study, as well as our recent studies in Kutch Basin, we conclude that the generic and specific diversity of orthophragminids decreases eastward from the peri-Mediterranean region to Indian subcontinent and to the western Pacific.  相似文献   

2.
Orthophragminids from the Bartonian Fulra Limestone in Kutch, India and the coeval units in Sulaiman Range in Pakistan suggest the establishment of a significant number of endemic species in the Indian subcontinent (Eastern Tethys). Among a total of fifteen species of Discocyclina, Orbitoclypeus and Asterocyclina, six of them appear to be confined to Indian subcontinent while seven species are common both to the peri-Mediterranean/Europe region (Western Tethys) and Indian subcontinent. Two species, Asterocyclina sireli, a four-ribbed species of possibly Indo-Pacific origin, and Orbitoclypeus haynesi that form large populations in Fulra Limestone, appear to have spread into North Africa and Turkey but not into European platforms as a response to Middle Eocene Climatic Optimum (MECO). The lack of Lutetian and Priabonian fauna in the studied sections, either due to a hiatus or unsuitable depositional environments, hampers the establishment of the actual stratigraphic ranges of the identified taxa. Our record provides us to characterize the orthophragminids in shallow benthic zone (SBZ) 17 for Eastern Tethys in detail by comparing the data from the above localities with those from the North Africa, Europe and Turkey, showing the change in diversity.  相似文献   

3.
Among the smaller foraminifers from the Lower Permian of Western Tethys (Pamir, Northern Afghanistan, Central and Eastern Iran, Armenia, and Turkey), foraminiferal assemblages characteristic of the Asselian, Sakmarian, Yakhtashian, Bolorian, and Kubergandian stages are distinguished. The first stratigraphic scheme based on smaller foraminifers is elaborated for the Lower Permian (Cisuralian Series) of Western Tethys. Eight biostratigraphic units distinguished in the Lower Permian and one in the Kubergandian Stage are ranked as beds with characteristic foraminiferal assemblages. At particular stratigraphic levels, the beds are recognizable in different paleogeographic provinces of the Tethyan Realm, which enables correlation between deposits concurrently accumulated under dissimilar climatic and facies conditions to be carried. Some of the distinguished beds are recognizable beyond the Tethyan Realm, for instance in the Donetsk basin, Cis-Urals, Pechora coal basin, and Spitsbergen. Among foraminifers that have been studied, 264 species and subspecies, including 16 new taxa, are identified. The following species and subspecies are identified and described for the first time: Hemigordius permicus beitepicus subsp. nov., H. pamiricus sp. nov., Neohemigordius afganicus sp. nov., N. carnicus sp. nov., N. bangi sp. nov., N. zulumarticus sp. nov., N. kubergandinicus sp. nov., Geinitzina grandella sp. nov., G. dentiformis sp. nov., G. bella sp. nov., Pachyphloia paraovata minima sp. nov., P. aucta sp. nov., Frondicularia porrecta sp. nov., Globivalvulina gigantea sp. nov., G. compacta sp. nov., and G. explicata sp. nov.  相似文献   

4.
A single confidently dated species of cephalopod is so far known in the Tremadocian of the southern Central Andean Basin (NW Argentina and southern Bolivia). This species belongs to the Eothinoceratidae and has a strong affinity mainly with Avalonia. During the Floian, a notable increase in diversity took place, with the appearance of a variety of families represented by several genera, in particular, within the Family Eothinoceratidae. In addition to the previously described species from southern Bolivia, we evaluate the other records of that family from the Central Andean Basin, and propose the following new taxa: Saloceras sikus sp. nov., Saloceras quena sp. nov., Mutveiceras gen. nov., and Mutveiceras cienagaensis sp. nov. We also describe Margaritoceras diploide, Margaritoceras sp., and Mutveiceras sp. From a palaeogeographic perspective, the cephalopod fauna shows affinities mainly with those of England, Wales, and the Montagne Noire (cold water Gondwana and peri‐Gondwana). As with other cephalopod faunas of mid to high palaeolatitudes, eothinoceratids occur along with other cephalopods forming assemblages of low morphological diversity. We interpret the forms described here as demersal with a subvertical poise, but capable of making rapid buoyancy changes, living in a wide spectrum of shallow offshore to shoreface settings. Copyright © 2014 John Wiley & Sons, Ltd.  相似文献   

5.
<正>One new genus and five new species of the family Evaniidae are described from the Early Cretaceous (Albian)Spanish amber of Penacerrada-Ⅰ(Province of Burgos),San Just and Arroyo de la Pascueta(both in the Province of Teruel):Cretevania alonsoi sp.nov.,C.montoyai sp.nov.,C.alcalai sp.nov.,C.rubusensis sp.nov.,and Iberoevania roblesi gen.and sp.nov.Taxonomic changes include Cretevania pristina(Zhang and Zhang,2000) comb.nov.,C.exquisita(Zhang,Rasmtsyn,Wang and Zhang,2007)comb.nov.,C.vesca(Zhang,Rasnitsyn,Wang and Zhang,2007)comb.nov.,and C.cyrtocerca(Deans,2004)comb.nov.,as a result of the reinterpretation of the genera Procretevania and Eovernevania.The new well preserved specimens of the genus Cretevania,together with the characters shown by the type specimens of the synonymized genera,give new information about their anatomical characters of taxonomical importance,and the genus Cretevania Rasnitsyn,1975 is re-diagnosed.The holotypes of the Russian species in amber have been revised.A cladistic analysis of fossil and extant groups of the superfamily Evanioidea is included.Cretevania had a wide palaeogeographic distribution,with the highest diversity known from Spain.The 13 known Cretevania species show a high interspecific variation mainly in wing characteristics,and a wide range of body and wing size.  相似文献   

6.
Three new species within the stigmaphronid genus TagsmiphronEngel and Grimaldi, 2009, and one new species within the megaspilid genus ConostigmusDahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage.  相似文献   

7.
The biodiversity record of graptolites from the Cheeseman's Creek Formation, considered herein as late Gisbornian (Caradoc) in age, has been substantially increased to fifteen taxa, including the new species Dicellograptus praemorrisi sp. nov. and Climacograptus vandenbergi sp. nov. Some of the records have global correlative significance enabling us to identify the wilsoni Biozone ( = calcaratus Biozone of eastern Australia). Several evolutionary lineages have been recognized:
  • 1 Dicellograptus moffatensis (Carruthers, 1858) → D. praemorrisi sp. nov → D. morrisi Hopkinson, 1871
  • 2 Glossograptus hincksi Hopkinson, 1872 → Glossograptus? sp.
  • 3 Climacograptus bicornis (J. Hall, 1847) → C. vandenbergi sp. nov. → C. lanceolatus VandenBerg, 1990.
Copyright © 2001 John Wiley & Sons, Ltd.  相似文献   

8.
The following new fossil trichopteran insects are described or revised from the Purbeck Limestone Group and Wealden Supergroup of southern England: Pteromixanum inviolatum gen. et sp. nov., P. ruderatum gen. et sp. nov., P. purbeckianum (Handlirsch) gen. et comb. nov. and P. poxwellense gen. et sp. nov. (Necrotauliidae);Purbimodus minor gen. et sp. nov., P. medius gen. et sp. nov., P. rasnitsyni gen. et sp. nov. and P. saxosus gen. et sp. nov. (Vitimotauliidae);Palaeoludus popovi gen. et sp. nov. (Dysoneuridae);Palaeotarsus desertus gen. et sp. nov. (Plectrotarsidae);Eucrunoecia ridicula gen. et sp. nov. (Lepidostomatidae);Palaeocentropus placidus gen. et sp. nov. (Calamoceratidae) and Helicophidae gen. et sp. incertae sedis. The general composition of the fauna is compared with Early Cretaceous faunas of Asia. The early appearance of several living families is noted.  相似文献   

9.
An Early Triassic (Griesbachian) gastropod fauna is reported from South China (Shanggan, Guangxi) and consists of four species: Bellerophon abrekensis, Wannerispira shangganensis Kaim & Nützel sp. nov., Naticopsis sp., and Palaeonarica guangxinensis Kaim & Nützel sp. nov. The taxon Wannerispira Kaim & Nützel nom. nov. replaces Pagodina Wanner non Van Beneden. This is the first report of Bellerophon abrekensis from China. Previously, it was only known from its type locality in Far East Russia. Wannerispira shangganensis sp. is the first certain Triassic report of the Permian subfamily Neilsoniinae and represents a holdover taxon. The neritimorph Palaeonarica is reported for the first time from the Early Triassic and this is the oldest occurrence of this genus. Compared with other Griesbachian gastropods, the present material is relatively well preserved so that the taxonomy rests on rather firm ground. Very few nominal taxa have been reported from the Griesbachian and therefore the present report presents substantial additional information about gastropods from the aftermath of the end-Permian mass extinction event. The gastropod association from Shanggan shares one species with Primorye, Far East Russia (B. abrekensis). Two species, W. shangganensis and P. guangxinensis, closely resemble specimens reported from the Griesbachian of Oman. This could suggest that Griesbachian gastropod faunas of the Tethys were rather homogenous although the data are still scarce.  相似文献   

10.
The mid-Cenomanian fish beds of Nammoura, Mont-Liban district, Lebanon contain a diverse fauna of aquatic and terrestrial vertebrates, a few crustaceans and moderately well-preserved plant remains of which a single species,Sapindopsis anhouryi , was previously described by Dilcher & Basson (1990). We add 11 species of ferns, gymnosperms and angiosperms of which Nammouria cretacea gen. et sp. nov, Nupharanthus cretacea gen. et sp. nov., Sapindopsis libanensis sp. nov. and Nammourophyllum altingioides gen. et sp. nov. are new taxa. The florule differs markedly from both Early Cretaceous and Turonian plant assemblages of the Middle East, thereby representing a distinct stage of the regional floristic evolution. Its phytogeographic affinities are with contemporaneous floras of North America, Central Europe and the Crimea. A combination of such features as xeromorphism, the prevalence of compound leaves, and the presence of deciduous angiosperm components and gymnosperms may indicate climatic conditions similar to those of the present day Mediterranean.  相似文献   

11.
<正>The forewing of a termite from Early Miocene lake sediments in Otago,southern New Zealand is figured and described.It exhibits the generic characters of the damp-wood termite Stolotermes Hagen,but differs from forewings of the known species in size and venation pattern and is described as Stolotermes kupe sp.nov.S.kupe represents the first confident record of fossil Stolotermitidae and extends the fossil record of the family back to the Early Miocene.It also is the first direct evidence of fossil Isoptera from New Zealand,though silicified termite faecal pellets,referable to Kalotermes brauni,have been previously described.S.kupe indicates that Stolotermitidae has been present in the Australasian region since at least the Early Miocene.  相似文献   

12.
A new genus with two new species, Orentalphila gravia gen. et sp. nov. and O. caloa sp. nov., and three new species in two known genera, Sinosciophila angustia sp. nov., Sinosciophila seboa sp. nov. and Similsciophila undulata sp. nov., are described from the Lower Cretaceous of Yixian Formation at Huangbanjigou, Beipiao City, western Liaoning, China. These new taxa, representing the first records of mesosciophilids from this locality, provide morphological information and diversity for Mesosciophilidae in the Early Cretaceous. An updated key to known genera of Mesosciophilidae is provided. In addition, according the generic diagnosis revised by Zhang, 2007, Mesoplecia antiqua Hao and Ren, 2009 should be transferred to Mesosciophila Rohdendorf, 1946.  相似文献   

13.
中国南方泥盆系发现颗石藻   总被引:1,自引:0,他引:1       下载免费PDF全文
颗石藻作为一种超微远洋浮游生物,出现于侏罗纪里阿斯世(Lias),繁盛于晚白垩世至第三纪。最近又有所进展。哈克(Hag,1981)面告,最老的颗石藻出现于石炭纪和二叠纪。本文所报道之颗石藻发现于晚泥盆世,是截至目前为止最老的颗石藻化石。 颗石藻演化迅速,分布广泛,形态特征明显,而且数量众多,对划分对比海相地层有一定的意义。本文只对研究区所采标本的颗石藻进行鉴定、描述和分类。  相似文献   

14.
Yanjiahella gen. nov., a distinctive fossil taxon from the Early Cambrian Yanjiahe Biota in Yichang, Hubei Province, is characterized by: 1) an overall body plan composed of a stalk and a calyx (also arms in some species); 2) a stalk with closely spaced, transverse wrinkles and a medial, longitudinal ridge; and 3) a calyx preserved with three-dimensional radiating striations, as well as a probable madreporite in some specimens. Three species that can be assigned to this genus, which are Y. ancarpa sp. nov., Y. monocarpa sp. nov., and Y. biscarpa sp. nov. Yanjiahella is comparable to certain echinoderm taxa in terms of morphology and some biological structures, which might indicate the early evolution and diversification of ancestral echinoderm lineages from the early Early Cambrian Yangtze Platform. The discovery of putative echinoderms in the Yanjiahe Biota provides significant clues about the origination and evolution of the deuterostome phylum Echinodermata.  相似文献   

15.
A Wordian (Middle Permian) cephalopod fauna consisting of four ammonoid species, Tauroceras aff. scrobiculatum (Gemmellaro), Agathiceras sp., Bamyaniceras orientale n. sp. and Pronoritidae gen. and sp. indet., and two nautiloid species, Tainoceras sp. and Orthocerida fam. indet., were recovered from the Bera South area, southern Pahang, Peninsular Malaysia. The outcrop represents the southernmost extent of the Bera Formation. Bamyaniceras and Tainoceras are recorded in Malaysia for the first time. The presence of T. aff. scrobiculatum suggests a regional correlation with an ammonoid-bearing bed of Sungai Cheroh, western Pahang, and permits global correlations with strata of northeastern Iraq, northern Oman and Sicily (Italy). A possible eastwards faunal migration through the southern Tethys is suggested.  相似文献   

16.
The occurrence of the genus Aquilapollenites in Upper Cretaceous and Neogene sediments of northwestern Pakistan is reported here. Aquilapollenites amplus, Aquilapollenites reductus, and Aquilapollenites sp. occur in the Maastrichtian palynomorph assemblage from an outcrop sample of the Mir Ali section, northern Waziristan. Aquilapollenites medeis in the Neogene Murgha Faqir Zai Formation of the Pishin Basin, Balochistan, is considered a reworked Cretaceous specimen. The Upper Cretaceous sediments of the Asian plate on the Tethys margin are considered to be the source of Aquilapollenites spp. in these samples.  相似文献   

17.
Sampling of Cenomanian fossil-rich horizons within the La Luna Formation of two localities in the Zulia and Trujillo states (northern Venezuela) yielded numerous shark teeth belonging to various species within the order Lamniformes (Mackerel sharks). Twelve lamniform species were identified including three new species (Squalicorax lalunaensis sp. nov., Squalicorax moodyi sp. nov., Acutalamna karsteni gen. et sp. nov.) and the genus Microcarcharias gen. nov. is proposed to accommodate with the peculiar morphology of the small-sized odontaspidid M. saskatchewanensis. Other taxa reported here include Cretoxyrhina mantelli, Cretolamna sp., cf. Nanocorax sp. and five Squalicorax species left in open nomenclature. This is the first report of chondrichthyans from the mid-Cretaceous of Venezuela and one of the few records of this group from the Cenomanian of South America. The composition of these assemblages suggests some degree of endemism in the La Luna Sea but also possible connexions with the Western Interior Seaway. One of the most striking features of these assemblages is the high anacoracid diversity (eight species) despite the corresponding outer shelf/upper slope palaeoenvironments of the La Luna Formation. The high diversity of these opportunistic predators is probably related to the high diversity of medium to large marine vertebrates that provided food resources.  相似文献   

18.
A well-preserved ammonoid fauna of Early Dienerian age has long been known from the lower portion of the Candelaria Formation in the old Candelaria silver mining district in Mineral and Esmeralda Counties, Nevada, but for a number of reasons, this fauna has never been studied in detail nor illustrated. Previous authors assigned this ammonoid fauna to the Early Dienerian Proptychites candidus Zone of Canada. In reality, it more closely resembles the Tethyan faunas than the higher palaeolatitude Canadian faunas, thus indicating the presence of some degree of equatorial faunal exchange between opposite sides of the Panthalassic Ocean during Early Dienerian time. It also indicates the onset of a provincialism, which contrasts with the cosmopolitan Griesbachian faunas. A rigorous taxonomic analysis of the Candelaria fauna allows us to differentiate the following ten species, which include two new species and one new genus (Mullericeras nov. gen.) belonging to the new family Mullericeratidae: Ambites lilangensis (Krafft, 1909), Ambites aff. radiatus (Brühwiler, Brayard, Bucher and Guodun, 2008), Ussuridiscus sp. indet., ??Koninckites?? aff. kraffti Spath, 1934, Mullericeras spitiense (Krafft, 1909), Mullericeras fergusoni nov. sp., Mullericeras sp. indet., Proptychites haydeni (Krafft, 1909), Proptychites pagei nov. sp., Vavilovites sp. indet. and Parahedenstroemia kiparisovae Shigeta and Zakharov, 2009. This Early Dienerian fauna correlates with the Ambites fauna known from the base of the Ceratite Marls in the Salt Range and from the base of the ??Meekoceras?? beds in Spiti (northern Gondwanian margin). The fauna also permits the precise dating of a shelfal anoxic episode on the equatorial North American margin. This anoxic event correlates in time with similar palaeoceanographic changes in the southern Tethys, which indicates that the Early Triassic biotic recovery was at least partly shaped by such discrete, short events rather than by pervasive and lingering adverse environmental conditions.  相似文献   

19.
The lower part of the Pawpaw Shale in northeast Texas yields an abundance of diminutive limonitic ammonites, the assemblage being: Tetragonites sp. juv., Engonoceras serpentinum (Cragin, 1900), Conlinites wrighti gen. et sp. nov., Cantabrigites wenoensis (Adkins, 1920), Neophlycticeras (Neophlycticeras) sp., Neophlycticeras (Paradolphia) occidentalis sp. nov., Stoliczkaia (Stoliczkaia) clavigera Neumayr, 1875, Stoliczkaia (Lamnayella) worthense (Adkins, 1920), Stoliczkaia (Shumarinaia) asiatica Matsumoto and Inoma, 1975, Enigmaticeras riceae gen. et sp. nov., Flickia simplex Pervinquière, 1907, Ficheuria pernoni Dubourdieu, 1953, Ficheuria americana sp. nov., Anisoceras armatum (J. Sowerby, 1817), Hamites venetzianus Pictet, 1847, Mariella (Mariella) worthensis (Adkins and Winton, 1920), Mariella (Mariella) asper sp. nov., Lechites (Lechites) comanchensis (Adkins, 1920), Worthoceras worthense (Adkins, 1920), and Scaphites hilli Adkins and Winton, 1920.This assemblage is correlated with the Mortoniceras (Subschloenbachia) rostratum Subzone of the European sequence on the basis of the occurrence of that species in the underlying Weno Limestone, in the passage beds between the Pawpaw Shale and the succeeding Main Street Limestone, and the lower part of the Main Street.Some previous authors have interpreted the diminutive limonitic ammonites of the Pawpaw as being “dwarf” or “stunted”. The overwhelming majority of individuals are nuclei of much larger ammonites, with smaller numbers of adults of small species that occur in other facies and geographic locations, together with a smaller number still of paedomorphic dwarfs. The fauna is interpreted as a preservational one, where diagenetic pyrite formed in relatively small voids, either the early phragmocone chambers of larger ammonites, or the phragmocone and body chambers of genuinely diminutive species which oxidised to limonite during the weathering process.  相似文献   

20.
Forty-seven samples from Upper Cretaceous sections penetrated by the Kachi-1 and Inga-1 wells in the South Yellow Sea Basin have been analysed for their spore and pollen content. Thirty-five species of 18 spore genera and 54 species of 28 pollen genera are documented. One new monotypic genus, Diporocolpopollenites, and its type species, D. kachiensis sp. nov., are erected, and Dilwynites Harris, 1965, and its type species, D. granulatus Harris, 1965, are emended. There are also three new combinations: Ephedripites eocaenicus (Selling, 1944), E. praeclarus (Chlonova, 1961), and Retitricolpites anguloluminosus (Anderson, 1960). Two palynological zones are erected: anAquilapollenites attenuatus Assemblage Zone, which encompasses deposits that are considered to be latest Campanian–Early Maastrichtian in age, and an Aquilapollenites eurypteronus Assemblage Zone for sections that have been dated as Late Maastrichtian. The assemblages are typical of the Yenisey-Amur Subprovince of the Aquilapollenites (floral) Province. Lowland floodplain to shallow, commonly mesotrophic, lacustrine environments of deposition are indicated. The climate was probably wet subtropical, with rainfall being somewhat higher during the Late Maastrichtian than through the latest Campanian–Early Maastrichtian.  相似文献   

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