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1.
The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments.  相似文献   

2.
A partial, associated skeleton with feather impressions of a bird from early Cretaceous (Neocomian) deposits in Mongolia constitutes a new family and order (Ambiortidae, Ambiortiformes). The specimen presents a mosaic of archaic and specialised characters within the Class Aves and shows that advanced carinate birds existed some 10 to 12 million years after Archaeopteryx, lending tentative support to the idea that Archaeopteryx may not be representative of the state of avian evolution in the late Jurassic. The new specimen and numerous feather impressions from other early Cretaceous localities in Mongolia and the Soviet Union, indicate that birds were probably common in early Cretaceous biotas.  相似文献   

3.
Austrotriconodon mckennai and Austrotriconodon sepulvedai, from the Campanian Los Alamitos Formation, Patagonia, Argentina were originally described as triconodont mammals and the sole members of the family Austrotriconodontidae. These mammals were represented by isolated cheek teeth originally regarded as molariforms, but their peculiar morphology later raised doubts about their purported triconodont affinities. Nevertheless, the morphological bases supporting the alternative taxonomic views have not been fully documented. We present here detailed comparisons of Austrotriconodon with other Late Cretaceous taxa and conclude that Austrotriconodon specimens should be assigned to Meridiolestida and Mesungulatoidea. These isolated teeth are likely premolars and might represent unknown dental positions of already described species or correspond to taxa that are yet to be formally recognized. According to our interpretation, there is still no record of Cretaceous triconodonts in South America, but we support the triconodont affinities for Jurassic taxa from the Cañadón Asfalto Formation in central Patagonia.  相似文献   

4.
Metriorhynchidae is a clade of marine-adapted crocodilians known from several Middle Jurassic–Early Cretaceous specimens collected predominantly in South America and Europe, but poorly known in the northern margin of Gondwana. The “Portomaggiore crocodile” is the most complete specimen of an Italian metriorhynchid to date: it consists of a partial skeleton that has been provisionally referred to an unnamed species of Late Jurassic Metriorhynchus or Geosaurus. The specimen is preserved in the reddish, nodular limestone of the Rosso Ammonitico Veronese Formation (Bajocian–Tithonian); new data on microfossil associations constrain the age of the metriorhynchid to the late Bajocian–earliest Bathonian. On the basis of cranial synapomorphies, the “Portomaggiore crocodile” falls as the closest sister-taxon of the Late Jurassic–Early Cretaceous geosaurines, and is referred to Neptunidraco ammoniticus gen. et sp. nov. It is unique among Middle Jurassic metriorhynchids in showing an incipient streamlining of the skull, shared with Late Jurassic and Cretaceous taxa. Since Neptunidraco is the oldest known member of Metriorhynchidae, its phylogenetic position supports the hypothesis that the timing of the initial metriorhynchid and geosaurine diversifications should start in the Bajocian.  相似文献   

5.
Here we describe an isolated tooth of a metriorhynchid crocodylomorph from the Hybla Formation (Aptian, Lower Cretaceous) of Rocca Chi Parra quarry (Montagna Grande, Calatafimi, Trapani Province), Sicily, Italy. The specimen shares with the Upper Jurassic taxon Plesiosuchus manselii a mediolaterally compressed conical tooth crown, noticeable lingual curvature, mesial and distal carinae with microscopic, rectangular contiguous denticles, strong distal curvature of the mesial margin, and the presence of weak 'carinal flanges' on the labial and lingual surfaces (which are preeminent at the mid-crown). This suite of morphologies is also present in an unnamed Valanginian (Lower Cretaceous) plesiosuchinan from France. However, the Sicilian tooth differs from these taxa in having more pronounced carinae, and faint apicobasally aligned enamel ridges. It also differs from P. manselii in having more extensive 'carinal flanges' on the labial surface. The specimen extends the known geological range of Metriorhynchidae and Thalattosuchia by approximately 7–8 million years. This overturns previous hypotheses of Metriorhynchidae becoming extinct early in the Early Cretaceous.  相似文献   

6.
Rare, isolated teeth of Corysodon multicristatus sp. nov. are described from two levels in the Atherfield Clay Formation (Early Aptian, Early Cretaceous) of Atherfield Point on the Isle of Wight, UK. Ten teeth of the new species were recovered from 1095 kg of washed and graded sediment residues. The teeth themselves are very small (around 0.5 mm high) and possess a distinctive crown bearing a tiered series of transverse crests adapted for rasping. Details of the dental architecture of the Atherfield Clay Formation specimens clearly indicate that the Cretaceous material differs significantly from the teeth of the type species for the genus, Corysodon cirinensis, recorded from the Kimmeridgian of northern France and Switzerland. C. multicristatus is the first substantiated record of the genus from the Early Cretaceous, thereby extending the stratigraphic range of the genus from the latest Jurassic, and the geographical range from continental Europe to the UK.  相似文献   

7.
To date three taxa of troodontid theropod dinosaurs have been recognized from Upper Cretaceous strata in two regions of the Kyzylkum Desert in Uzbekistan. The Cenomanian Khodzhakul Formation in the southwestern Kyzylkum Desert has yielded isolated serrated teeth and some postcranial bones of an indeterminate troodontid. In the central Kyzylkum Desert troodontids are known from the Cenomanian Dzharakuduk Formation (Urbacodon itemirensis) and the Turonian Bissekty Formation (Urbacodon sp.). Urbacodon itemirensis is known from a single dentary whereas Urbacodon sp. is represented by isolated teeth, maxilla and dentary fragments, a partial braincase, and some postcranial bones. The troodontid affinities of Urbacodon are supported by several synapomorphies: presence of a subotic recess; reduced basal tubera placed directly under the occipital condyle; maxilla participating in the margin of the external naris; nutrient foramina on dentary situated within a deep lateral groove; dentary without distinct interdental plates; large number of small dentary and maxillary teeth; teeth constricted between root and crown; anterior dentary teeth smaller, more numerous, more closely spaced than those in the middle of the tooth row, and implanted in a groove; posterior dorsal vertebrae with tall and posterodorsally tapering neural spines; and presence of a midline sulcus on the neural arches of distal caudals. Among Troodontidae, Urbacodon resembles Byronosaurus, Gobivenator, and Xixiasaurus in the absence of serrations on the tooth crowns and having premaxillary teeth that are D-shaped in cross-section. However, phylogenetic analysis did not recover a clade of Asiatic troodontids with unserrated teeth.  相似文献   

8.
Late Jurassic and Early Cretaceous charophyte assemblages from the northern part of the Aquitaine Basin in south-west France are reviewed here to understand their palaeoecological, palaeobiogeographical and biostratigraphic features. Three sites were studied: the Tithonian-lower Berriasian of Chassiron, and the Berriasian of Cherves-de-Cognac and Angeac-Charente. Abundant porocharaceans, less abundant clavatoraceans and scarce characeans recorded in Cherves-de-Cognac and Angeac-Charente indicate that brackish water environments were substituted by freshwater environments eastwards. The occurrence of Clavator grovesii var. grovesii and morphotypes intermediate with C. grovesii var. discordis in the same areas is significant from a biostratigraphic viewpoint, since these species belong to the Maillardii, Incrassatus and Nurrensis European charophyte biozones, representing the Berriasian. This observation refutes a previous dating of the Angeac-Charente site and highlights the absence of Hauterivian–Barremian records in northern Aquitaine, which is in contrast to the more complete Lower Cretaceous record in southern Aquitaine. These contrasting records could be due to differences in the available sedimentary space produced by the opening of the Bay of Biscay during the Barremian.  相似文献   

9.
10.
While pterosaurs occur in the Lower Jurassic strata of Britain and Germany, only the family Rhamphorhynchidae is currently known found in both. A newly discovered humerus from the Whitby Mudstone Formation of Lincolnshire challenges this and is distinguished from all other Lower Jurassic British pterosaurs by its possession of a quadrangular deltopectoral crest. This is a rare morphotype which only occurs in Eudimorphodon, Austriadraco, Raeticodactylus, Carniadactylus and Campylognathoides. The Lincolnshire humerus compares well with these taxa but is identified as a cf. Campylognathoides sp. based on its age and palaeobiogeography. The genus Campylognathoides is a common pterosaur in the Toarcian Posidonia Shale of Germany and the new humerus supports continuity of pterosaur populations across central Laurasia.  相似文献   

11.
12.
Extensive new material of scalpelliform cirripedes, including articulated individuals, from the Late Jurassic (Tithonian) Kimmeridge Clay of Dorset, southern England, is described and used as a basis for taxonomic revision of the family Zeugmatolepadidae. Two subfamilies, the Zeugmatolepadinae and Martillepadinae nov., are established and two new genera (Martillepas, Concinnalepas) in the latter subfamily are described from the Upper Jurassic and one (Icenilepas) from the Upper Cretaceous Chalk. Material from the Kimmeridge Clay of Dorset also includes the oldest fossil representative of the family Pollicipedidae for which a new genus and species, Etcheslepas durotrigensis, are erected, and the oldest calanticid, Cretiscalpellum sp. nov. Jurassic taxa established by T.H. Withers in 1928, on the basis of sparse material, are redescribed, and referred to the new genera as Martillepas ovalis, M. costata, M. hollisi, Concinnalepas concinna and Etcheslepas fragilis.  相似文献   

13.
In Mexico, the Upper Jurassic to lowermost Cretaceous La Casita and coeval La Caja and La Pimienta formations are well-known for their abundant and well-preserved marine vertebrates and invertebrates. The latter include conspicuous inoceramid bivalves of the genus Anopaea not formally described previously from Mexico. Anopaea bassei (Lecolle de Cantú, 1967), Anopaea cf. stoliczkai (Holdhaus, 1913), Anopaea cf. callistoensis Crame and Kelly, 1995 and Anopaea sp. are rare constituents in distinctive Tithonian–lower Berriasian levels of the La Caja Formation and one Tithonian horizon of the La Pimienta Formation. Anopaea bassei was previously documented from the Tithonian of central Mexico and Cuba, while most other members of Anopaea described here are only known from southern high latitudes. The Mexican assemblage also includes taxa which closely resemble Anopaea stoliczkai from the Tithonian of India, Indonesia and the Antarctic Peninsula, and Anopaea callistoensis from the late Tithonian to ?early Berriasian of the Antarctic Peninsula. Our new data expand the palaeogeographical distribution of the high latitude Anopaea to the Gulf of Mexico region and substantiate faunal exchange, in the Late Jurassic–earliest Cretaceous, between Mexico and the Antarctic Realm.  相似文献   

14.
Yabeinosaurus was the first lizard genus described from the Early Cretaceous Jehol Biota of China. The holotype of the type species, Y. tenuis, is an immature skeleton but it has been lost for decades. A second species, Y. youngi, was erected based on another immature skeleton and is distinguished by its longer, more gracile limbs. In 2001, a juvenile skeleton from the Jingangshan locality of Liaoning was designated as the neotype for Y. tenuis. Subsequently, several further specimens from other localities, juvenile and adult, have been attributed to this species. Here we describe a second lizard specimen from the neotype locality. In anterior tooth shape, finger-like mandibular angular process, cranial sculpture, maxilla shape, and short, robust humerus, the new specimen resembles adult and subadult material currently attributed to Y. tenuis, but it differs in having bicuspid posterior teeth and a straight rather than a hooked angular process of the mandible. We propose a new species (Y. bicuspidens sp. nov.) for this specimen which represents the first occurrence of bicuspid teeth in an Early Cretaceous lizard from China. However, the recognition of a second species at the neotype locality raises a taxonomic problem. Due to the immaturity of the designated holotype and neotype of Y. tenuis, they cannot be coded for the jaw and dental characters that distinguish the two robust-limbed species. This renders Yabeinosaurus tenuis a nomen dubium. Here we propose that Yabeinosaurus tenuis should be treated as a historic taxon, permitting retention of the generic name under ICZN rules, and erect a second new species (Y. robustus sp. nov.) for the principal robust-limbed morphotype in the Jehol Biota with monocuspid posterior teeth and a hooked angular process.  相似文献   

15.
Lepisosteid fishes are well known from the Upper Cretaceous of Europe, but only by fragmentary remains from some Cenomanian and Campanian–Maastrichtian deposits. Here we report various cranial and postcranial remains of gars, discovered in the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút (Bakony Mountains, Hungary). These remains represent one of the most diverse assemblages of lepisosteid fish material from Upper Cretaceous continental deposits of Europe. Based on tooth morphology, scale-microstructure and the features of the supracleithrum we refer these remains to the genus Atractosteus. Besides some uncertain remains from the Cenomanian of France and Spain, the Santonian aged fossils from Iharkút represent the oldest undisputable occurrence of the family Lepisosteidae in the European continental Cretaceous. Using tooth crown morphology, the surface microstructure of the ganoid scales and the anatomy of the supracleithrum a review of the Late Cretaceous lepisosteid record suggests the occurrence of both Atractosteus and Lepisosteus in the European archipelago.  相似文献   

16.
Diverse thoracican cirripedes from the Hauterivian of the Hannover district of northern Germany are described, including seven species, belonging to five genera. Of these, a new genus belonging to the Scalpellidae, Jaegerscalpellum, includes one Hauterivian species, J. elegans sp. nov., an Aptian species, J. comptum (Withers, 1910) and an Albian species, J. politum (Darwin, 1851) are also referred to it. A new Cretiscalpellum, C. mutterlosei sp. nov. is described from the Hauterivian, and C. matrioni sp. nov. is described from the Middle Albian of France. The oldest record of the Unilatera Gale, 2018, Pedupycnolepas pulcher sp. nov. is described from the Hauterivian; this displays typical shell structure of the group, retained by living Verrucidae. Finally, four species of Zeugmatolepadidae, subfamily Martillepadinae, are recorded from the Hauterivian, including Martillepas hausmanni (Koch and Dunker, 1836), M. decoratus sp. nov., M. auriculum sp. nov. and Etcheslaepas borealis (Collins, 1990). The Hauterivian fauna from Hannover shows affinities both with Late Jurassic and later Cretaceous (Aptian-Cenomanian) forms, and includes the earliest scalpellids, unilateran (Pedupycnolepas) and Cretiscalpellum species known. It constrains the age of the Cretaceous cirripede evolutionary radiation to the earliest Cretaceous.  相似文献   

17.
A new species of hybodontid shark is described for the Lower Cretaceous of Brazil. The type specimen is derived from pre-Aptian strata of Malhada Vermelha Formation, Lima Campos Basin (Ceará State, northeastern Brazil), with referred material from the Missão Velha Formation of Araripe Basin, northeastern Brazil. The new taxon differs from other Planohybodus species by the presence of a stronger tooth ornamentation characterized by simple, usually non-branching cristae that nearly reach the apex of the main cusp as well as distinctly divergent lateral cusplets. In addition, certain North-American specimens formerly attributed to Hybodus are identified as Planohybodus. The new species presented herein, in addition to the North-American remains, represents an important contribution to the knowledge of the distribution of Planohybodus, expanding the geographic range of the genus to South and North America in the Late Jurassic and Early Cretaceous.  相似文献   

18.
Rapid radiation of the Bittacidae during the Jurassic resulted in high diversity within this family of Mecoptera. More than 40 species within 23 genera have been described from this period. However, around the end of the Jurassic the abundance and diversity of Bittacidae decreased and only seven species in five genera are known to date from the Cretaceous. A new specimen from the basal Cretaceous of England, Tytthobittacus jarzembowski sp. nov., described here, represents the eighth species and the second fossil representative of this family from the European Cretaceous. This hangingfly belongs to an extant and relictual genus previously known only from Australia. The paper also includes a review of all known Cretaceous bittacids and a re-examination of European species Antiquanabittacus nanus Petrulevičius and Jarzembowski, 2004.  相似文献   

19.
Recent work in the Tropic Shale by the Museum of Northern Arizona reveals a high biodiversity for plesiosaurs along the western margin of the Cretaceous Western Interior Seaway during the Cenomanian and the Turonian. This paper describes a new species of polycotylid plesiosaur from the Tropic Shale, Dolichorhynchops tropicensis, which adds to the known biodiversity from this time period. The identification is based on two specimens, a well-preserved, nearly complete skeleton including the skull and an additional specimen with only fragmentary skeletal elements. The material shares several synapomorphic characters with Dolichorhynchops, including the shape of the temporal fenestrae, the shape of the sagittal crest, the trend of the ectopterygoids, the morphology of the teeth, and the number of teeth within the mandibular symphysis. D. tropicensis differs from other species of Dolichorhynchops in a greater size range of the teeth, moderately constricted dorsal vertebrae, angled anterior processes on the coracoids, and the presence of well-defined facets on the propodials. The specimens are of Early Turonian age (based primarily on ammonite biostratigraphy and bentonite marker beds), and their discovery extends the known stratigraphic range for Dolichorhynchops back by approximately seven million years. This suggests that Dolichorhynchops tropicensis and Trinacromerum bentonianum, a closely related polycotylid, co-existed in the Western Interior Seaway.  相似文献   

20.
In the Murihiku Terrane of New Zealand, U-Pb detrital zircon ages in Murihiku Supergroup sandstones of Late Triassic, Jurassic and possibly earliest Cretaceous age have a marked youngest age component that is close to, and sometimes coincident with, established biostratigraphic ages, thus reflecting contemporary volcanism. However, youngest Huriwai Group samples yield 137–142 Ma zircon age components (earliest Early Cretaceous) in conflict with palynofloras that suggest only a latest Jurassic age. This is resolved if the age of the Jurassic/Cretaceous boundary is lowered to ca. 140 Ma. Older, reworked zircons are mainly Early Jurassic, Late Triassic and Late Permian reflecting an enduring exhumed magmatic arc source nearby. This might be in the adjacent Median Batholith but as a Murihiku sediment source its Jurassic, Triassic and Permian elements are not well-matched in terms of extent, age and bulk compositions. A connection between the Murihiku (proximal forearc) and Waipapa Composite (distal accretionary wedge) terranes is probable, with a common magmatic arc, speculatively situated in the New England Orogen, eastern Australia.  相似文献   

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