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At hatching the larvae of flatfish closely resemble the bilateral symmetric larvae of other teleosts, especially perciforms. Literature data show that transformation to asymmetric benthic juveniles normally occurs at body lengths between 10 and 25 mm. Unexpectedly, minimal size at its completion (including eye migration) can be 4.1 mm SL and maximal size is over 72 mm. In this paper we consider the functional requirements for a successful switch from a symmetric pelagic larva to a typical asymmetric juvenile benthic flatfish partly based on evidence from other teleosts. The unfavourable period of eye migration and transition to a benthic habitat requires some food reserves and rewiring and/or recalibration of vision and gravity-associated structures utilised previously by the still symmetric larvae for e.g. food detection. Binocular fixation of the prey probably occurs in that stage. Critical or sensitive periods occurring during development of fish larvae suggest that a completely functional symmetric stage of development must precede transformation. The normal size range in flatfish larvae at transformation seems to confirm our considerations. Recent data on temperature effects during development provide an explanation for metamorphosis at the minimal size. Some evidence for paedomorphic heterochrony in flatfish larvae is presented. 相似文献
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虾肤诱食剂改善南美白对虾植物蛋白日粮的作用 总被引:6,自引:0,他引:6
用900尾平均体质量为0.106 g/尾±0.012 g/尾的南美白对虾(Peneausvannamei)进行虾肽改善植物蛋白日粮试验,结果表明:(1)大豆粕、杂粕组南美白对虾的成活率显著高于鱼粉组(P<0.05);(2)大豆粕、鱼粉组南美白对虾相对生长率、饲料系数、虾体组成相似,杂粕组最差,且与大豆粕、鱼粉组有显著性差异(P<0.05)。结果表明,在南美白对虾日粮中虾肽作为营养性诱食剂可显著改善植物蛋白的适口性,提高南美白对虾对大豆蛋白的的消化吸收功能,达到与鱼粉组相同的养殖效果,从而明显降低南美白对虾日粮成本。 相似文献
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LIU Luofu HUO Hong GUO Yongqiang CHEN Lixin LI Shuangwen ZHAO Yande LI Yan WANG Ping CHEN Zhijun 《中国地球化学学报》2008,27(1):1-8
The results presented in this paper indicated that the carbazole-type compounds have high thermal stability and also show stability in oxidation and bio-degradation. This kind of compounds still has high concentrations and a complete distribution in the analyzed dry asphalt samples, showing that they are particularly useful in the study of hydrocarbon migration of the paleo-pools. The difference in the contents of nitrogen compounds in the Silurian dry asphalts from the Awati, Tabei and Tazhong areas is attributed to the difference in the extent of oxidation and (or) bio-degradation for the areas; the Awati and Tabei areas underwent relatively strong oxidation and bio-degradation. During the first stage of hydrocarbon pool formation in the Silurian system in the Tazhong and Tabei areas of the Tarim Basin (at the end of the Silurian period) and at the second stage in the Awati area (in Permian), the hydrocarbons experienced a long-distance migration. 相似文献
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Petroleum Migration Direction of the Silurian Paleo-pools in the Tarim Basin, Northwest China 总被引:1,自引:0,他引:1
LIU Luofu ZHAO Ye HUO Hong CHEN Lixin CHEN Yuanzhuang ZHAO Suping LI Chao LI Shuangwen GUO Yongqiang LI Yan 《《地质学报》英文版》2008,82(1):174-183
The results obtained in this paper indicate that carbazole-type compounds have high thermal stability and also show stability in oxidation and bio-degradation. This kind of compounds still has a high concentration and complete distribution in the analyzed dry asphalt samples, showing that they are particularly useful in studying petroleum migration of paleo-pool. During the basin's first-stage of oil-gas pool formation in the Silurian in Tazhong and Tabei areas of Tarim Basin (at the end of Silurian period) and the second-stage in the Awati area (in Permian), the petroleum experienced a long-distance migration. During the formation of the Silurian paleo-pools in Tazhong Uplift at the end of Silurian, the petroleum mainly came from the lower and middle Cambrian source rocks in the Manjiaer sag. The petroleum migrated towards the southwest-south entering the Silurian reservoir beds in Tazhong first. Then, it further migrated within Silurian from northwest to southeast along the highs of the Structural Belts to the region of the Silurian pinchout boundary in Tazhong. In Tabei Uplift, during the first-stage of pool formation, the petroleum was also from the lower and middle Cambrian source rocks in the Manjiaer sag. It migrated northwest entering the Silurian reservoir beds in the Tabei Uplift firstly, and then the migration continued in the same direction within the Silurian reservoirs and finally the petroleum was trapped in higher positions. During the second-stage pool formation in the Silurian beds in the areas around Awati sag, the petroleum mainly came from the lower-middle Cambrian source rocks in the Awati sag. The petroleum migrated from the generation center to Silurian reservoirs in all directions around the sag through major paths, and the petroleum was finally trapped in higher locations. 相似文献
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Subtle traps or oil pools have become an important exploration play in the Dongying Depression, Bohai Bay Basin, east China. Despite recent successes in exploration, the formation mechanisms of subtle traps are still not well understood. The majority of subtle oil pools in the Dongying Depression are developed in the middle interval of the Es3 Member of the Paleogene Shahejie Formation with the subtle traps being primarily of lenticular basin-floor turbidite sands encompassed in mudstones. Oil in the subtle traps was previously thought to have migrated directly from the surrounding source rocks of the same formation (Es3). Detailed geochemical investigation of 41 oils and 41 rock samples from the depression now indicates that the oils from the subtle traps cannot be correlated well with the surrounding Es3 source rocks, which are characterized by high Pr/ Ph (〉1), low Gammacerane/C30hopane, representing a freshwater lacustrine setting. In contrast the oils features low Pr/Ph (〈1) and relatively high Gammacerane content, showing a genetic affinity with the underlying Es4 source rocks, which also have the same qualities, indicating a brackish lacustrine setting. Oils in the Es3 subtle traps are probably derived from mixed sources with the contribution from the upper Es4 source rocks predominating. Therefore unconventional oil migration and accumulation mechanisms need to be invoked to explain the pooling of oils from the ES4 source rocks, which probably came through a thick low interval of the Es3 source rocks with no apparent structural or stratigraphic pathways. We suggest that the subtle oil migration pathway probably plays an important role here. This finding may have significant implications for future exploration and the remaining resource evaluation in the Dongying Depression. 相似文献