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1.
We review the previously described Late Cretaceous (Santonian) bird remains from the Csehbánya Formation in the Bakony Mountains of Hungary, augmenting initial work by Ősi ( 2008 ), and add a number of newly collected fossils. All together, the eight fossil specimens so far collected from this site are important to our understanding of avian evolution because they document a large range of taxon body sizes from at least one major lineage (Enantiornithes) and come from a critically undersampled time period in the Cretaceous. Globally, very little fossil bird material has been collected from the middle stages of the Late Cretaceous, the Coniacian and Santonian; most known taxa are either Early Cretaceous (ca. 120 Ma) in age or are from the terminal Campanian and Maastrichtian (ca. 70–65 Ma). Indeed, one of the Csehbánya Formation fossil birds is recognized as a new taxon of large enantiornithine, an avisaurid apparently similar in its largely unfused foot morphology to the Argentine Soroavisaurus and to the North American Avisaurus. The Central European records reviewed in this paper highlight the wide distribution of some Late Cretaceous fossil birds, particularly avisaurid enantiornithines, and lead us to a brief discussion of avian biogeography at the end of the Mesozoic. Copyright © 2010 John Wiley & Sons, Ltd.  相似文献   

2.
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3.
Although body fossils of shorebirds and shorebird-like species are extremely rare from the Cretaceous, rapid increase in the discovery of bird footprint sites provides valuable alternate evidence to help fill gaps in the story of the early evolution of shorebird-like species. Newly discovered bird tracks from the Albian-Cenomanian Dakota Formation in northeastern Utah represent the first report of the ichnogenus Koreanaornis from North America and only the second report of bird tracks from this formation. These tracks are not attributable to Aquatilavipes as previously claimed. Three well-preserved trackways are described and provisionally referred to Koreanaornis cf. hamanensis (Kim). However, a review of the ichnotaxonomy of shorebird ichnites reveals that this ichnotaxon also closely resembles the Miocene ichnospecies Avipeda sirin (Vyalov). This latter comparison points to the need for a thorough evaluation of the similarity between Mesozoic and Cenozoic avian ichnotaxa, which may be over-split in some cases and under-differentiated in others.The new material helps distinguish ichnogenus Koreanaornis from the larger bird track Aquatilavipes, which is more abundant and widespread in North America. In some cases Aquatilavipes has been incorrectly used as a catch-all ichnogenus both in North America and Asia. The Dakota Formation stratigraphy at the tracksite indicates that the track makers lived in a marginal marine paleoenvironment. However, despite the widespread distribution of such facies, often replete with dinosaur tracks, the bird track record of the Dakota Formation, and the Cretaceous of the western USA remains relatively sparse in comparison with other areas such as east Asia.  相似文献   

4.
The Fairpoint Member of the Fox Hills Formation (upper Maastrichtian) in Meade County, South Dakota, USA, contains an osteichthyan assemblage indicative of transitional to marine shoreface deposits. The fauna consists of: Lepisosteus sp., Paralbula casei, Cylindracanthus cf. C. ornatus, Enchodus gladiolus, Hadrodus sp., and indeterminate osteichthyans with probable affinities to the Siluriformes and Beryciformes. The Fairpoint fauna is of limited species diversity and in this character mirrors many other Upper Cretaceous North American osteichthyan assemblages. Comparison to Upper Cretaceous chondrichthyan diversity and consideration of the structure of Cretaceous marine food webs suggest that osteichthyans are strongly under-represented in the Upper Cretaceous of North America. The small size and poor preservation potential of many Upper Cretaceous North American osteichthyans probably account for much of this observed paucity. Fairpoint osteichthyans are members of families that survive the Cretaceous–Paleocene boundary extinction event. Some of these genera and families are still extant and occur in a wide array of modern fresh, brackish, and shallow marine environments.  相似文献   

5.
The occurrence of the genus Aquilapollenites in Upper Cretaceous and Neogene sediments of northwestern Pakistan is reported here. Aquilapollenites amplus, Aquilapollenites reductus, and Aquilapollenites sp. occur in the Maastrichtian palynomorph assemblage from an outcrop sample of the Mir Ali section, northern Waziristan. Aquilapollenites medeis in the Neogene Murgha Faqir Zai Formation of the Pishin Basin, Balochistan, is considered a reworked Cretaceous specimen. The Upper Cretaceous sediments of the Asian plate on the Tethys margin are considered to be the source of Aquilapollenites spp. in these samples.  相似文献   

6.
The first specimen of Aturoidea to be recorded in East Asia has been found in the Upper Cretaceous Sada limestone in Shimanto City, Kochi Prefecture, Japan. The specimen is one of the few representatives from the Upper Cretaceous, along with species known from Libya, Angola, and India. The specimen is very similar to A. mathewsonni from the Paleocene deposits in California. However, we describe the specimen as A. cf. mathewsonni, as it slightly differs from A. mathewsonni in the shape of the lateral lobe of the suture. The finding reveals that Aturoidea had already lived in waters around East Asia in the Late Cretaceous.  相似文献   

7.
The Lower Cretaceous Mural Limestone marks the maximum marine incursion into southeast Arizona during Aptian-Albian time and records the middle Cretaceous transition from coral-dominated to rudist-bivalve dominated reefs. Upper Mural Limestone facies are most often dominated by corals. However, rudists form significant frameworks at some localities, one of which is described in this paper. The paleoenvironmental distribution of three potential reef-builders (corals, rudists, and ‘oysters’) were studied at this patch reef locality. Corals built the framework of the inner reef core. The rudist Petalodontia initially gained a foothold in sheltered areas among corals and subsequently built a framework in the outer reef core. Caprinid rudists formed mounds in the outer reef to back reef areas. The rudists Toucasia and Monopleura and the oyster-like bivalve Chondrodonta formed beds or were scattered in the reef-flank and shelf lagoon sediments and did not contribute to the reef framework.Upper Mural Limestone reefs are important examples of the coexistence of corals and rudists during this middle Cretaceous faunal transition period. This study supports the idea that rudist-bivalves initially colonized protected back-reef areas early in the Cretaceous and only later in the Cretaceous did rudists dominate reef frameworks.  相似文献   

8.
The isotopic and geochemical studies of the Upper Cretaceous-Cenozoic flysch sequences of the Kamchatka Peninsula and southern Koryak region revealed that they were formed at least from two sources: one depleted (T) with low 87Sr/86Sr and high positive SrNd(T) values and one enriched (T) with high 87Sr/86Sr and negative SrNd(T) values. The enriched source was likely represented by complexes of ancient upper continental crust. The subduction-related rocks and, to a lesser extent, basalts of mid-oceanic ridges or back-arc basins could serve as a juvenile source for most of the flysch sediments. The Upper Cretaceous flysch sediments differ from their Cenozoic analogues in composition. The Upper Cretaceous rocks are dominated by enriched upper crustal material. The Cenozoic sequences of the Ukelayat Trough and Paleocene-Eocene sequences of the Kumroch Range contain a substantial amount of island-arc volcanoclastic material; the Eocene flysch of Karaginskii Island is compositionally similar to the Upper Cretaceous flysch sequences.  相似文献   

9.
Lepisosteid fishes are well known from the Upper Cretaceous of Europe, but only by fragmentary remains from some Cenomanian and Campanian–Maastrichtian deposits. Here we report various cranial and postcranial remains of gars, discovered in the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút (Bakony Mountains, Hungary). These remains represent one of the most diverse assemblages of lepisosteid fish material from Upper Cretaceous continental deposits of Europe. Based on tooth morphology, scale-microstructure and the features of the supracleithrum we refer these remains to the genus Atractosteus. Besides some uncertain remains from the Cenomanian of France and Spain, the Santonian aged fossils from Iharkút represent the oldest undisputable occurrence of the family Lepisosteidae in the European continental Cretaceous. Using tooth crown morphology, the surface microstructure of the ganoid scales and the anatomy of the supracleithrum a review of the Late Cretaceous lepisosteid record suggests the occurrence of both Atractosteus and Lepisosteus in the European archipelago.  相似文献   

10.
The Family Afrograptidae is a ‘conchostracan’ group with multiple radial costae reaching to the umbo on their carapaces. It comprises four described genera: Afrograpta, Camerunograpta, Congestheriella and Graptoestheriella with a total of thirteen described species which are occasionally reported from the Jurassic and the Cretaceous in Africa, Europe and South America (i.e. Afrograpta from the Upper Cretaceous of Cameroon; Camerunograpta from the Jurassic to Cretaceous of Cameroon; Congestheriella from the Jurassic to Upper Cretaceous of the Congo Basin, Brazil, Bulgaria, Venezuela and Argentina; and Graptoestheriella from the Upper Jurassic to Lower Cretaceous of Brazil). A new genus and a new species, Surreyestheria ockleyensis gen. et sp. nov., belonging to the Family Afrograptidae from the Lower Cretaceous (lower Barremian) Upper Weald Clay Formation of Ockley Village, Surrey County, southern England is described in this paper. The new genus mainly differs from the other four genera by the special reticulate ornamentation on its carapace. It indicates that the Family Afrograptidae was more diverse and more widely distributed in the late Mesozoic than previously supposed. Afrograptidae is a special branch of Estheriellina the latter originating in the late Palaeozoic and the former in the early Mesozoic. Afrograptids, as a whole had been widespread across Pangea in the Early Jurassic.  相似文献   

11.
Three Schizolepis species collected from the Lower Cretaceous layer of the Huolinhe Basin,Inner Mongolia,China are described.These fossils are Schizolepis longipetiolus Xu XH et Sun BN sp.nov.,which is a new species,Schizolepis cf.heilongjiangensis Zheng et Zhang,and Schizolepis neimengensis Deng.The new species is a well-preserved female cone,slender and cylindrical in shape.The seed–scale complexes have long petioles and are arranged on the cone axis loosely and helically.The seed scales are divided into two lobes from the base.Each lobe is semicircular or elongate ligulate in shape,widest at the middle or the lower middle part,with an obtuse or bluntly pointed apex.The inner margin is almost straight and the outer margin is strongly arched.On the surface of the lobe,there are longitudinal and somewhat radial striations from the base to the margin.The seed is borne on the adaxial surface at the base or middle of each lobe.Schizolepis was established in 1847,and,although more than twenty species have been discovered and reported,its phylogenetic position is controversial because of the imperfection of fossils.Most authors have considered there to be a close evolutionary relationship between Schizolepis and extant Pinaceae.Here,we analyze characteristics and compare Schizolepis with Picea crassifolia Kom,which is morphologically most similar to Schizolepis.The results indicate that the genus probably has a distant evolutionary relationship with extant Pinaceae.A detailed statistical analysis of the global paleogeographic distribution of Schizolepis showed that all the fossils of this genus appeared in strata ranging from the Upper Triassic to the Lower Cretaceous in the North Hemisphere,being rare in the Upper Triassic and Lower Jurassic,but being very common from the Middle Jurassic to the Lower Cretaceous,and particularly abundant in the Lower Cretaceous.According to the statistical results,we speculate that the genus originated in Europe in the Late Triassic then spread from Europe to Asia between the Late Triassic and the Late Jurassic.In the Early Cretaceous most species existed in China’s three northeastern Provinces and the Inner Mongolia Autonomous Region and adjacent areas.Combining the paleogeographic distribution of the genus with ancient climatic factors,we deduced that Schizolepis began to decline and became extinct in the Early Cretaceous,and the reason for its extinction is closely related to the icehouse climate during the Early Cretaceous.  相似文献   

12.
We report on a new enantiornithine bird, Dunhuangia cuii, gen. et sp. nov., from the Lower Cretaceous Xiagou Formation of the Changma Basin, northwestern China. Although the material is incomplete, Dunhuangia cuii preserves unique coracoidal and sternal morphologies that distinguish it from other known enantiornithines; this specimen represents only the second enantiornithine reported from the Changma Basin that is diagnostic at the species level. This study enriches our understanding of the enantiornithine component of this ornithuromorph-dominated, Early Cretaceous avifauna.  相似文献   

13.
Summary Integration of new mineral chemical, geochronological and structural data from the Texel Complex yielded information on (re)crystallization and deformation processes in metapelites, eclogites and tonalitic orthogneisses during eclogite facies metamorphism. Maximum P–T conditions reached 1.2 to 1.4 GPa and 540–620 °C in the Upper Cretaceous. In tonalitic orthogneisses and metapelites, substantial garnet growth took place prior to eclogite facies metamorphism and Sm–Nd data indicate the presence of pre-Cretaceous mineral relics. In contrast, complex garnet-growth and -resorption processes are inferred for eclogites, which produced characteristic atoll microstructures and occurred close to the pressure peak of a single, coherent high pressure event. Garnet Sm–Nd data indicate eclogite facies crystallization at 85 ± 5 Ma. While eclogites retained information on the maximum burial stage, matrix phases in metapelites and orthogneisses were intensely recrystallized during the amphibolite facies metamorphic decompression. All the meso- and macro-scale deformation structures formed during the high pressure event and subsequent exhumation. The major mylonitic foliation is represented by the high pressure phases but was refolded during amphibolite facies exhumation. A biotite-whole-rock Rb–Sr age of 70–80 Ma indicates that cooling below about 300 °C occurred in the Upper Cretaceous. Supplementary material to this paper is available in electronic form at Appendix available as electronic supplementary material  相似文献   

14.
A re-examination of fossil material from the Late Cretaceous Cambridge Greensand Member (CGM) of the West Melbury Marly Chalk Formation revealed a number of new specimens of edentulous pterosaur jaw fragments previously identified as shark fin spines and fish jaws and accessioned under the epithet ‘cestraciontid finray’ and ‘jaws of fish’. These are now recognised as pterosaurian jaw tips and referred to Ornithostoma sedgwicki Seeley, 1891 and Azhdarchoidea indet. This material increases the diversity of edentulous pterosaurs from the CGM.The edentulous pterosaur Ornithostoma sedgwicki Seeley, 1891 from the Cretaceous Cambridge Greensand of eastern England is reviewed. The holotype specimen is confirmed as a fragment of a premaxilla/maxilla of a non-tapejarid azhdarchoid on account of the conspicuous curvature of the dorsal and occlusal margins posteriorly and the presence of small neural foramina on the lateral margins. Neural foramina are not seen on jaws of members of the Pteranodontia, a group to which O. sedgwicki was included previously. The referral of O. sedgwicki to Azhdarchoidea eliminates the single known Lower Cretaceous occurrence of Pteranodontidae, restricting the temporal range of this taxon to the Upper Cretaceous. Postcranial material referred to O. sedgwicki from the type horizon is regarded as indeterminate Pterosauria.  相似文献   

15.
Three new species within the stigmaphronid genus TagsmiphronEngel and Grimaldi, 2009, and one new species within the megaspilid genus ConostigmusDahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage.  相似文献   

16.
The Chikkim Formation as exposed in the Tethyan Himalaya (India) has been studied at its type locality, using planktonic foraminifera for a detailed biostratigraphic elaboration. Divided into two members, the Lower and Upper Chikkim members, this formation ranges in age from Albian to early Maastrichtian(?), and reaches a maximum thickness of 150 m. Examination of thin sections has yielded 34 species of foraminifera in five genus-level assemblages. The Lower Chikkim Member is about 55 m thick; its basal portion is of Albian age based on the presence of Biticinella breggiensis and Planomalina buxtorfi. At 26 m above the base, Whiteinella archaeocretacaea documents OAE 2 (Oceanic Anoxic Event 2), and thus the Cenomanian/Turonian boundary in this section. The carbonate sequence is capped by a Santonian-age hardground with iron oxide crusts and bioturbation. Macrofossils, including belemnites (at the base) and irregular echinoids (upper part), are present. The basal carbonaceous marls of the Upper Chikkim Member yield both large (benthic) rotaliid as well as planktonic foraminifera (Globotruncanita elevata, Gl. stuartiformis, Gl. stuarti, Gansserina gansseri and others), indicating a Campanian age. The co-occurrence of Gl. elevata and G. gansseri in a single thin section results either from condensation or reworking in the basal part of the Upper Chikkim Member. Late Cretaceous index foraminifera such as Gl. elevata document deposition within the Tethyan Realm. The original thickness of the Upper Chikkim Member is uncertain, but would have been around 100 m; the unit appears markedly reduced through weathering at a height of about 5000 m above sea level. Equivalent sediments are exposed in the Zanskar area to the northwest, and in Nepal and Tibet. Cretaceous Oceanic Red Beds (CORBs) are probably missing due to the proximality of these pelagic settings.  相似文献   

17.
We describe a large collection of fossil ‘waterfowl’ bones that are referable to the extinct clade Presbyornithidae (Anseriformes). All of these fossils were collected between 1971 and 1994 from Upper Paleocene and Lower Eocene sediments at the Tsagaan Khushuu site in the Gobi Desert of southern Mongolia. The collection includes specimens referred to a new small species within the genus Presbyornis Wetmore, 1926 as well as large numbers of bones that we place in the genus Presbyornis. On this basis of the Tsagaan Khushuu collection we suggest that several species of Presbyornis likely coexisted in this region; indeed, the presence of large numbers of middle‐sized, morphologically consistent but probably ecologically disparate species at the Tsagaan Khushuu site is consistent with the range of variation seen, for example, in taxa of extant dabbling ducks (Anatini). Although the anatomy and phylogenetic position of Presbyornithidae (in particular Presbyornis) are well known, this material from Mongolia further demonstrates the prevalence of these birds in aquatic and semi‐aquatic habitats by the earliest Paleogene. Because presbyornithids are also well documented from the late Cretaceous, their palaeoecology and morphological diversity provides a clue to selective avian survivorship across the Cretaceous‐Paleogene (K‐Pg) boundary. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

18.
The western part of the Tarim Basin in Xinjiang is one of the main areas in China where the marineCretaceous is well developed. The Upper Cretaceous Yingjisha Group represented mainly by sediments of lit-toral, near-shore neritic and estuarine facies is divided in ascending order into the Kukebai Formation, theOytak Formation, the Ygezya Formation and the Tuylouk Formation. For about thirty years, the basal beds of the Kukebai Formation had been considered to be the lowermostmarine horizon of the Cretaceous in the western part of the Tarim Basin, which represents the earliest trans-gression of the Cretaceous Sea into this region. Recently. marine trace fossils, Ophiomorpha nodosa, O.tuberosa and Thalassinoides? spp. were found in abundance and fine preservation from the upper subcycle andupper part of the lower subcycle of the Kezlesu Group underlying the Kukebai Formation. The fact indicatesthat the marine transgression there took place earlier than the Kukebaian. Process of transgression and regression and change of environment in the West Tarim Basin during the pe-riod from the late Early Cretaceous to the end of the Cretaceous is also discussed in this paper.  相似文献   

19.
The Jurassic paleogeographic position of the Pontides is not well studied because of insufficient paleomagnetic data. For this reason, a paleomagnetic study was carried out in order to constrain the paleolatitudinal drift of the Turkish blocks during the Jurassic period. A total of 32 sites were sampled from volcanic and volcanoclastic rocks of the Lower/Middle Jurassic Kelkit formation (Eastern Pontides), Mudurnu formation (Sakarya continent) and Upper Jurassic–Lower Cretaceous Ferhatkaya formation exposed around Amasya region (Eastern Pontides). Rock magnetic experiments demonstrate that the main ferromagnetic mineral is pseudo-single-domain titanomagnetite in these rocks. Paleomagnetic analysis revealed two main components of the natural remanent magnetization during stepwise thermal and alternating field demagnetization. The first component is a low-coercivity (unblocking temperature) component with a direction sometimes similar to that of the earth’s present field or a viscous component. The second component, which is interpreted as the characteristic remanent magnetization (ChRM) direction, has low to high coercivity properties between 20 and 100 mT or unblocking temperatures between 300 and 580°C. A positive fold test at the 95% level of confidence proved that the ChRM of the sites is primary. Paleomagnetic directions calculated for the Kelkit formation in the Eastern Pontides have a mean direction of D = 334.8°, I = 49.7°, α 95 = 7.1° after tilt-correction. A mean direction of D = 332.2°, I = 48.5°, α 95 = 14.6° was obtained from the volcanoclastic rocks of the Mudurnu formation, and D = 324.3°, I = 43.3°, α 95 = 9.5° was calculated for the Upper Jurassic–Lower Cretaceous limestones/Ferhatkaya formation of the Amasya region. The Jurassic rocks in the Eastern Pontides and Mudurnu region are considered to represent products of the rifted Neo-Tethys ocean, while the Upper Jurassic–Lower Cretaceous sediments in Amasya are related to basin-filling materials. The data suggest that the Kelkit formation was formed at 30.5°N paleolatitude and the equivalent Mudurnu formation at 29.5°N paleolatitude. The paleolatitude of the Eastern Pontides indicates that this rifting block was separated from Eurasia by a marginal basin instead of being a part of Eurasia. The lower paleolatitude of the Amasya region at 24.8°N in the Upper Jurassic to Lower Cretaceous clearly indicates southward drift of the Turkish blocks during the Jurassic to Lower Cretaceous period together with the motion of Eurasia.  相似文献   

20.
Three taxa of Late Cretaceous aquatic angiosperms, Queruexia angulata （Lesq.） Krysht., Cobbania corrugata （Lesq.） Stockey et al. and Nelumbites cf. extenuinervis Upchurch et al. from Jiayin of Heilongjiang, NE China, are described in detail. Among them, Cobbania and Nelumbites from the Upper Cretaceous in China are reported for the first time. The aquatic angiosperm assemblage of Queruexia-Cobbania-Nelumbites appears to imply a seasonal, warm and moist environment in the Jiayin area during the Santonian-Campanian time.  相似文献   

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