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1.
A re-examination of fossil material from the Late Cretaceous Cambridge Greensand Member (CGM) of the West Melbury Marly Chalk Formation revealed a number of new specimens of edentulous pterosaur jaw fragments previously identified as shark fin spines and fish jaws and accessioned under the epithet ‘cestraciontid finray’ and ‘jaws of fish’. These are now recognised as pterosaurian jaw tips and referred to Ornithostoma sedgwicki Seeley, 1891 and Azhdarchoidea indet. This material increases the diversity of edentulous pterosaurs from the CGM.The edentulous pterosaur Ornithostoma sedgwicki Seeley, 1891 from the Cretaceous Cambridge Greensand of eastern England is reviewed. The holotype specimen is confirmed as a fragment of a premaxilla/maxilla of a non-tapejarid azhdarchoid on account of the conspicuous curvature of the dorsal and occlusal margins posteriorly and the presence of small neural foramina on the lateral margins. Neural foramina are not seen on jaws of members of the Pteranodontia, a group to which O. sedgwicki was included previously. The referral of O. sedgwicki to Azhdarchoidea eliminates the single known Lower Cretaceous occurrence of Pteranodontidae, restricting the temporal range of this taxon to the Upper Cretaceous. Postcranial material referred to O. sedgwicki from the type horizon is regarded as indeterminate Pterosauria.  相似文献   

2.
Cretornis hlavaci Frič, 1881 from the Upper Cretaceous (Turonian) of Czech Republic is a valid taxon referred to Azhdarchoidea based on having a saddle-shaped humeral head, pneumatic foramen on proximal humerus present on anterior side and absent on posterior side, elongate deltopectoral crest with subparallel proximal and distal margins, pneumatic foramen absent on distal side of humerus, metacarpals I–III not articulated with carpus and displaced on anterodorsal side of wing metacarpal, and wing metacarpal much longer than humerus. Absence of a pneumatic foramen on posterior side of proximal humerus suggests attribution of Cretornis hlavaci to Neoazhdarchia. It has a unique construction of the distal ulna with a dorsal articulation surface placed distinctly proximal to the tuberculum shared only with the non-azhdarchid azhdarchoid Montanazhdarcho minor from the Campanian of North America. Cretornis hlavaci differs from the latter taxon by the structure of its humerus and distinctly longer wing metacarpal. It is more derived than “Tapejaridae” but shares with Azhdarchidae the deltopectoral crest of the humerus displaced distally from humeral head. Cretornis cannot be assigned to Azhdarchidae because of the oval cross section of the second wing phalanx. A unique rhombic outline of the distal humerus of Cretornis hlavaci is a possible autapomorphy for this taxon. Its wing span estimated as 1.5–1.6 m. This is the first taxon of non-azhdarchid pterosaurs known from the Upper Cretaceous of the Eastern Hemisphere.  相似文献   

3.
Four isolated cervical vertebrae from the Kimmeridge Clay Formation (Upper Jurassic, Kimmeridgian) of Abingdon, Oxfordshire, England are identified as from a pliosaurid plesiosaurian sauropterygian on account of their shortness relative to width and height, their near platycoelous nature and the location of tall rib facets on the centrum body. They are noteworthy for their size, with a maximum width of 269 mm, maximum height of 222 mm and maximum length of 103 mm. Simple scaling and comparisons with cervical vertebrae of Mid Jurassic pliosaurs Peloneustes and Liopleurodon, and the Early Cretaceous Stenorhynchosaurus and Sachicasaurus suggest a total body length of between ~ 9.8 m and 14.4 m for the Abingdon Kimmeridge Clay pliosaur. Likely the true length was towards the higher end of this range.A genus and species cannot be confidently determined on the basis of the described material, but they likely belong to Pliosaurus sp. or a similar animal, for which a precise neck length is not known. We estimate a neck length of 0.77 m for Pliosaurus ?brachyspondylus based on the average cervical lengths provided for specimen CAMSM J.35991.  相似文献   

4.
An isolated jaw fragment from the Late Cretaceous (Cenomanian) Cambridge Greensand Member of the West Melbury Marly Chalk Formation previously identified as a cestraciontid shark fin spine is referred to the pterosaur clade Azhdarchoidea on account of its lateral and occlusal foramina and edentuly. The specimen differs from the azhdarchoid Ornithostoma sedgwicki from the same deposit in having flat lateral surfaces and an acute dorsal/ventral apex. The specimen is similar in overall morphology to CAMSM B40085 from the same horizon and probably represents the corresponding jaw but from a different individual. Likely these specimens represent a new taxon but are considered too fragmentary to diagnose at present. A remarkably similar and distinctive morphology is found in unnamed pterosaur jaws from the Kem Kem Group (?Albian-Cenomanian) of Morocco, supporting the idea of faunal similarity between these two distant localities.  相似文献   

5.
A new specimen of pterosaurian metacarpal IV from the Early Cretaceous (Berriasian) Durlston Formation of Dorset, southern England, closely resembles those of the Chinese dsungaripterid Dsungaripterus weii Young, 1964 and the Central Asian azhdarchid Azhdarcho lancicollis Nessov, 1984. However, the new specimen lacks the thickened bony wall typical of Dsungaripteridae and is therefore regarded as azhdarchoid. An mcIV with a markedly offset distal condyle with well-developed diaphyseal constriction behind the distal condyle and asymmetrical condylar margins may be an autapomorphy of Dsungaripteridae + Azhdarchoidea, while the same condition with bone thickening may typify Dsungaripteridae. The new specimen also provides osteological evidence supporting claims for large wing-spanned pterosaurs in the Berriasian of southern England based previously only on ichnological finds. The new specimen suggests a wingspan of between 2.9 and 3.7 m and represents the largest pterosaur from the Berriasian.  相似文献   

6.
《Cretaceous Research》2012,33(6):774-780
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

7.
In contrast to the Barremian Wessex Formation on the Isle of Wight, the remains of small theropods are rare in the Berriasian-Valanginian Hastings Group of the English mainland. Both units are part of the dinosaur-rich Wealden Supergroup (Berriasian-Aptian) of southern Britain. Here we report the cervical vertebra of a small dinosaur from the Pevensey Pit at Ashdown Brickworks, a site located northwest of Bexhill, East Sussex. The pit yields a rich assemblage of vertebrate fossils from the Valanginian Wadhurst Clay Formation of the Hastings Group. The new specimen, a near-complete but water-worn posterior cervical vertebra, is tiny (total centrum length = 7.1 mm) but evidently from an adult theropod. Its large hypapophysis, X-shaped neural arch and amphicoelous centrum suggest referral to Maniraptora, and the subparallel anterior and posterior articular surfaces imply that it does not belong to a deinonychosaur. The X-shaped neural arch recalls a similar condition seen in oviraptorosaurs while the high neural canal/articular surface ratio (0.70) is bird-like. The specimen is significant in representing the first maniraptoran to be reported from the Hastings Group but is otherwise indeterminate. By comparing the specimen to better known maniraptorans and estimating the proportions of the animal to which it belongs, we suggest that the total skeletal length of this maniraptoran was somewhere between 16 and 40 cm. It may therefore have been among the smallest of known Mesozoic dinosaurs.  相似文献   

8.
A new long-necked sauropod dinosaur, Yunmenglong ruyangensis gen. et sp. nov., is erected on the basis of an incomplete skeleton from the late Early Cretaceous Haoling Formation of the Ruyang Basin, Henan Province. The characters of the anterior cervical vertebrae, the shape of the neural canal of the dorsal vertebra and the ball-shaped distal end of the neural spine of caudal vertebrae with coarse surfaces differ from other long-necked sauropod dinosaurs. The new genus has characters in common with both Euhelopus and Erketu; it represents the first long-necked sauropod dinosaur recorded from central China to date. The rod-like, well-developed epipophyses and the pleurocoels on the cervical vertebrae indicate that it may be close to Euhelopus, an observation also confirmed by a phylogenetic analysis, which shows that Erketu, Yunmenglong and Qiaowanlong form a clade, and are more derived than Euhelopus.  相似文献   

9.
The Upper Greensand Formation, in part mainly underlain by the Gault Formation and overlain by the Chalk Group, has extensive cliff outcrops in the Dorset and East Devon Coast World Heritage Site (WHS). The argillaceous Gault, up to 20 m thick in the Isle of Purbeck, is poorly exposed due to its involvement in extensive landslides, but the exposures of Upper Greensand are the most complete in England. The Gault (Middle Albian) rests unconformably on progressively older Jurassic and Triassic strata when traced westwards and becomes more arenaceous in the same direction. On the east Devon coast, the Upper Greensand comprises up to 55 m of sandstones and calcarenites that were deposited in fully marine, shallow-water environments. The formation is divided into three members there (Foxmould, Whitecliff Chert and Bindon Sandstone) each bounded by a prominent erosion surface. The full thickness of the Upper Greensand, up to 60 m, was formerly exposed in cliffs in the Isle of Purbeck in and adjacent to the steeply dipping limb of the Purbeck Monocline. The lower (Foxmould) part of the succession is similar to that in east Devon, but the upper part (White Nothe Member) is lithologically different and probably the correlative of only the Bindon Sandstone. Much of the fauna of the Gault and Upper Greensand of the WHS is not age-diagnostic with the result that the ages of parts of the succession are still poorly known. However, diverse ammonite assemblages recorded from a few thin beds in the lower and highest parts of the succession show that all except one of the Albian ammonite zones is present.  相似文献   

10.
Sections in the Gault and of the Gault-Woburn Sands junction exposed in the Leighton Buzzard area of Bedfordshire are described. These give a much clearer picture of the ammonite zonal stratigraphy than hitherto. The depositional history of the Albian sediments is discussed. The relationship of the Shenley Limestone to the regularis nodule beds in the south of the area is demonstrated, together with the nature of the mixed regularis and kitchini nodule beds seen in the central area. Periods of erosion occurred in post-kitchini mammillatum Zone times, and after renewed sedimentation in the eodentatus Subzone. Overlying these beds are clays of spathi Subzone age which pass laterally into glauconitic marginal loams against a platform of Woburn Sands in the northern part of the area. Upon this platform occur knolls of bedded Silty Beds, capped by Shenley Limestone, against which sediments of high spathi, intermedius and niobe Subzones age thin. The base of the Upper Gault, of high cristatum Subzone age, rests non-sequentially upon an eroded surface of the Lower Gault. Clays of orbignyi Subzone age are overlain by a bed of phosphatic nodules representing the lower part of the varicosum Subzone, which in turn is overlain by thick clays which may in part be of varicosum and auritus Subzones age.  相似文献   

11.
A small to medium-sized pterodactyloid pterosaur (wingspan approximately 1.10 m) from the Upper Jurassic (middle-late Tithonian) marine deposits of the Vaca Muerta Formation of Patagonia (Los Catutos area, central Neuquén Province, Argentina) is reported. The specimen lacks the skull but constitutes a nearly complete postcranial skeleton, which includes cervical and dorsal vertebrae; a few thoracic ribs; both pectoral girdles; the left pelvic girdle; a proximal right wing (humerus, ulna, and radius) and metacarpal IV; a left wing that lacks only wing phalanx four; and both hindlimbs, the right one without the foot. Ontogenetic features suggest that the new fossil corresponds to a relatively mature individual, probably a subadult. Observed characters support its assignment to the Archaeopteroactyloidea, a basal clade within the Pterodactyloidea. This specimen is the second pterosaur from Los Catutos and the most complete Jurassic pterosaur so far known from South America.  相似文献   

12.
A new somphospondylan sauropod (Dinosauria, Titanosauriformes): Gannansaurus sinensis gen. et sp. nov. from the Late Cretaceous Nanxiong Formation of Ganzhou Basin, Jiangxi Province of southern China is erected based on one nearly complete dorsal vertebra and a mid-caudal vertebra. It is characterized by two centroprezygapophyseal lamina fossae; large, square-shaped concavity formed by centroprezygapophyseal lamina and dorsal margin of the centrum; large infraparapophyseal fossa; three openings present within the large lateral concavity, which occupy 65% length of the centrum; the posterior centroparapophyseal lamina (PCPL) and the anterior centroparapophyseal lamina (ACPL) weakly developed; the posterior centrodiapophyseal lamina (PCDL) separated by an infradiapophyseal fossa into two branches dorsally; parapophyseal and diapophyseal laminae cross to form "K" configuration. It shares some characters with the Early Cretaceous form-Euhelopus, indicating that it is more closely related to Euhelopus rather than to other titanosauriform sauropods. This means Gannansaurus may have a close phylogenetic relationship with Euhelopus.  相似文献   

13.
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

14.
Early Cretaceous sediments of Aptian–Albian age outcrop at Munday’s Hill Quarry, Bedfordshire, England. Previous papers describing the section have resulted in different terminologies being applied. The Lower Cretaceous in Bedfordshire is represented by sediments belonging to the Lower Greensand Group and the Gault Clay Formation. Within the Lower Greensand Group in the study area the Woburn Sands Formation, are of Aptian–Albian age. Selected samples have been analysed for palynology. The analysis reveals diverse palynomorph assemblages, including well-preserved dinoflagellate cysts and sporomorphs. Comparison of the assemblages with published records indicates that the lower samples are of Late Aptian age. Forms recorded include common Kiokansium unituberculatum, Cerbia tabulata, Aptea polymorpha and Cyclonephelium inconspicuum. An Early Albian age is indicated for the uppermost sample.  相似文献   

15.
A new dinosaur sauropod from the Golmayo Formation (upper Hauterivian-lower Barremian) in Soria province (Spain) is described as Soriatitan golmayensis gen. et sp. nov. The new material consists of one tooth, three dorsal vertebrae with ribs, a partial sacrum, five caudal centra, two caudal vertebral spines, one chevron, a humerus, an ulna, a radius, two partial ilia, two ischia, a fragment of pubis, and a partial femur. Cladistic analysis identified the material as belonging to Brachiosaruidae within Titanosauriformes. Sauropod material from the Golmayo Formation shares a diagnostic feature with Abydosaurus, Cedarosaurus, Tastavinsaurus and Venenosaurus including anteriorly deflected anterior–middle caudal neural spines and with Cedarosaurus a proximodistally straight lateral margin between the proximal head and the shaft of the humerus. Eight characters are potential autapomorphies in the sauropod from Spain, including the presence of a large rectangular ventral ridge below the preacetabular process of the ilium, the same length of the pubic and ischiadic blades in the ischium, and two lateral ridges in the lateral area of the deltopectoral crest of the humerus. The presence of Early Cretaceous brachiosarids in both, North America and Europe, give support to the hypothesis of a connection between the tectonic plates of these continents at some point during the Early Cretaceous.  相似文献   

16.
The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments.  相似文献   

17.
Mongolian Paleontological Center (MPC) 100/1305 is one of the most complete ankylosaurid skeletons ever collected, and includes much of the postcranial skeleton and numerous in situ osteoderms. This specimen has been referred to Saichania chulsanensis, based on the similarity of the skull compared to the holotype of Saichania, MPC 100/151. However, MPC 100/1305 does not include a skull, and so referral of MPC 100/1305 to Saichania must be based on postcranial characters. Comparison of the postcrania of MPC 100/1305 and MPC 100/151 reveals several differences in the scapula, humerus, and metacarpals, indicating that MPC 100/1305 should not be referred to Saichania. Additionally, although it was previously reported that MPC 100/1305 was collected from the Baruungoyot Formation at Khulsan in Mongolia, collection records indicate that this specimen was instead collected from the Djadokhta Formation at Zamyn Khond. Two ankylosaurid species are known from the Djadokhta Formation of Mongolia and China, Pinacosaurus grangeri and P. mephistocephalus. There are no diagnostic characters in MPC 100/1305 that can be used to refer this specimen to P. grangeri or P. mephistocephalus, and there are a few differences between MPC 100/1305 and Pinacosaurus, such as the number of caudal vertebrae, and morphology of the coracoid, which have uncertain taxonomic implications. At present, MPC 100/1305 is best referred to Ankylosauridae indet., or cf. Pinacosaurus, based on its generally congruent morphology with Pinacosaurus and its provenance from the Djadokhta Formation, in which Pinacosaurus is the only recognized ankylosaurid taxon.  相似文献   

18.
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19.
We present a large, fragmentary skull and the humerus of a mosasaur (Squamata, Mosasauroidea) recovered from upper Maastrichtian beds of the López de Bertodano Formation in Marambio (=Seymour) Island, Antarctica. The material belongs to a large, adult individual with marked heterodonty as well as unusual humeral features. Different phylogenetic analyses returned the studied specimen within the Tylosaurinae, while the distinctive features of the skull and humerus allow distinguish it from the unique Antarctic known tylosaurine species, Taniwhasaurus antarcticus (Novas et al., 2002), as well as from other known Late Cretaceous mosasaurids from the Southern Hemisphere, thus, justifying the erection of a new taxon, Kaikaifilu hervei gen. et. sp. nov. The different dental types documented in the specimen studied have been previously recorded through isolated teeth from the same locality and were subsequently referred to several genera. This new find is relevant for assessing the previously known fragmentary records of Antarctic mosasaurids, suggesting that its local diversity could be more reduced than previously interpreted. The new material represents the youngest occurrence of tylosaurines in Antarctica.  相似文献   

20.
While pterosaurs occur in the Lower Jurassic strata of Britain and Germany, only the family Rhamphorhynchidae is currently known found in both. A newly discovered humerus from the Whitby Mudstone Formation of Lincolnshire challenges this and is distinguished from all other Lower Jurassic British pterosaurs by its possession of a quadrangular deltopectoral crest. This is a rare morphotype which only occurs in Eudimorphodon, Austriadraco, Raeticodactylus, Carniadactylus and Campylognathoides. The Lincolnshire humerus compares well with these taxa but is identified as a cf. Campylognathoides sp. based on its age and palaeobiogeography. The genus Campylognathoides is a common pterosaur in the Toarcian Posidonia Shale of Germany and the new humerus supports continuity of pterosaur populations across central Laurasia.  相似文献   

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