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1.
An incomplete forefin in the collections of the National Museum of Wales, Cardiff, is herein assigned to the rare leptonectid species, Leptonectes solei, known only from the west Dorset coast. It is only the third specimen of the species and is smaller than both the holotype and referred specimen. The new specimen is from the Lower Jurassic (lower Pliensbachian, Charmouth Mudstone Formation, Tragophylloceras ibex Ammonite Zone, Acanthopleuroceras valdani Ammonite Subzone) of Seatown, Dorset, UK. It confirms that all three species of Leptonectes were present in the Pliensbachian and expands the known diversity of ichthyosaurs for that time period. We show that isolated forefins of Leptonectes can be assigned to a species with a high degree of confidence.  相似文献   

2.
New coelacanth material from the Middle Triassic Prosanto Formation of the Ducan and Landwasser area near Davos in eastern Switzerland, Canton Graubünden, is described. A sub-complete individual is visible in ventral view, and shows details of its branchial apparatus. In particular, it possesses relatively large teeth on the ceratobranchials, and possible ossified hypobranchials. Few diagnostic characters are observable, and most of them are visible on the mandibles preserved in lateral view. This specimen shares characters with Ticinepomis peyeri, a smaller form from the Middle Triassic of Monte San Giorgio, whose holotype is re-described in part here. A second specimen, a fragmentary caudal skeleton shows the typical supplementary lobe of coelacanths, and meristic characters indicating probable close affinities with T. peyeri. We refer this material to Ticinepomis cf. T. peyeri. Because the new specimen is larger than the holotype, we refute the possible juvenile status of the small specimen from Monte San Giorgio. The new material of Ticinepomis from Canton Graubünden shows anatomical features not preserved on the holotype and allows the addition of new characters to a previously published data matrix of actinistians. A phylogenetic analysis is performed, which supports that Ticinepomis is nested among the Latimeriidae. The diversity of post-Palaeozoic coelacanths is assessed. The taxic diversity of observed occurrences shows a peak in the Early Triassic and a peak in the Late Jurassic, as detected in previous studies. When ghost lineages are included in the computation, the Late Jurassic peak is smoothened. By comparing the taxic diversity curves with the curve of average ghost lineage duration, we conclude that the Early Triassic peak of diversity was probably caused by a biological radiation, whereas the Late Jurassic peak of observed diversity is probably the result of a Lagerstätten effect.  相似文献   

3.
《China Geology》2021,4(2):197-204
The authors reassessed the taxonomic distinction of Iteravis huchzermeyeri and Gansus zheni, which are two species of Ornithuromorpha based on specimens from the same locality in western Liaoning and derive from the Jehol Biota. The detailed comparisons of the holotype and referred specimens of both species, reveal no anatomical features that distinguish these taxa as separate species. Some minor differences are considered to relate to ontogenetic or interspecific differences. The stratigraphic occurrence for both specimens is the Lower Cretaceous Jiufotang Formation. Accordingly, the authors conclude that Iteravis huchzermeyeri has priority, by 15 days, for this taxon and that Gansus zheni is a junior synonym. The diagnosis of Iteravis huchzermeyeri is revised based on further study on all specimens referred to this species. Its generic distinction from Gansus is maintained thereby removing a potential genus-level correlation linking Xiagou Formation in Gansu Province with the Jiufotang Formation in Liaoning Province.  相似文献   

4.
Late Albian ammonite faunas from the Aitamir Formation of the Koppeh Dagh Basin in northeast Iran are described and illustrated. These comprise 14 taxa, several of which are recorded from Iran for the first time, namely Anahoplites planus (formerly recorded from central Iran in open nomenclature only), Semenoviceras solidus, Epihoplites (Metaclavites) iphitus, Hysteroceras orbignyi and Pseudhelicoceras robertianum. New records of Placenticeras grossouvrei extend the stratigraphic range of this species downwards into the Late Albian; previously it was known from the Early and Middle Cenomanian only. The record of the rare E. (M.) iphitus fills a palaeobiogeographic gap between Crimea and Tajikistan, and the holotype of Spath is re-illustrated here. Additionally, Epihoplites trapezoidalis, from the Late Albian of Tajikistan, is relegated into the synonymy of Spath's species. A large number of taxa typical of the Late Albian (upper part of the Gault Clay Formation) of northwest Europe indicate close palaeobiogeographic affinities with the Koppeh Dagh Basin and faunal exchange across the Russian Platform and Transcaspia. The stratigraphic succession of the ammonite faunas is used for a biostratigraphic subdivision of the upper Aitamir Formation.  相似文献   

5.
The fossil clam shrimp genus Xibeiestheria Wang in Shen et al., 1982, a component of the well-known Early Cretaceous Yanjiestheria fauna, was originally described from the lower Aptian Jingchuan Formation of Yanchi County, Ningxia Hui Autonomous Region of northwestern China. The morphological re-examination under an SEM of the holotype of the type species Xibeiestheria pora Wang in Shen et al., 1982 has revealed new taxonomic features not previously recognized. These include: (1) growth lines with serrated lower margins, having a row of small pores; (2) widely spaced radial lirae intercalated cross bars on growth bands in the dorsal part of the carapace; (3) fine reticulum occurring within coarse reticulation on growth bands in the middle part of the carapace; (4) puncta radially aligned along the widely spaced radial lirae on growth bands in the lower part of the carapace.  相似文献   

6.
7.
A re-examination of fossil material from the Late Cretaceous Cambridge Greensand Member (CGM) of the West Melbury Marly Chalk Formation revealed a number of new specimens of edentulous pterosaur jaw fragments previously identified as shark fin spines and fish jaws and accessioned under the epithet ‘cestraciontid finray’ and ‘jaws of fish’. These are now recognised as pterosaurian jaw tips and referred to Ornithostoma sedgwicki Seeley, 1891 and Azhdarchoidea indet. This material increases the diversity of edentulous pterosaurs from the CGM.The edentulous pterosaur Ornithostoma sedgwicki Seeley, 1891 from the Cretaceous Cambridge Greensand of eastern England is reviewed. The holotype specimen is confirmed as a fragment of a premaxilla/maxilla of a non-tapejarid azhdarchoid on account of the conspicuous curvature of the dorsal and occlusal margins posteriorly and the presence of small neural foramina on the lateral margins. Neural foramina are not seen on jaws of members of the Pteranodontia, a group to which O. sedgwicki was included previously. The referral of O. sedgwicki to Azhdarchoidea eliminates the single known Lower Cretaceous occurrence of Pteranodontidae, restricting the temporal range of this taxon to the Upper Cretaceous. Postcranial material referred to O. sedgwicki from the type horizon is regarded as indeterminate Pterosauria.  相似文献   

8.
The results of Barnum Brown's 1937 expedition to the Almond Formation of Wyoming consisted of two unidentified ceratopsian skulls and a partial hadrosaurid specimen (AMNH 3651). The hadrosaurid is here attributed to the Maastrichtian genus Saurolophus, verifying previous biostratigraphic correlations of this formation using ammonite zones. Fossiliferous lower Maastrichtian formations occurring latitudinally between those of Alberta, Canada, and southwestern Texas, USA, such as the Almond Formation, are essential for testing the effects and duration of apparent hadrosaurid faunal segregation earlier in the Campanian, and indirectly aiding in the placement of faunal boundaries that are currently unknown for the late Campanian. The discovery of Saurolophus in Wyoming, a close relative of the Campanian genus Prosaurolophus, affirms that the segregation of hadrosaurid faunas established in the late Campanian (~75 Ma) continued for at least 3 million years. Combining occurrences of Saurolophus from Mongolia and the Moreno Formation of California with those of Alberta, Canada, this genus appears to have had one of the largest geographic ranges of any equivalent clade of hadrosaurid dinosaur, although species level distributions are still uncertain.  相似文献   

9.
The systematics of elasmosaurids are not well resolved partially because of the scarcity of well-preserved skull material. Among Weddellian elasmosaurids, one exception to this is the holotype of Tuarangisaurus keyesi from upper Campanian–lower Maastrichtian levels of the Maungataniwha Sandstone Member of the Tahora Formation, Mangahouanga Stream, inland Hawke's Bay, New Zealand. The material preserves an almost complete cranium and mandible. This material is re-described here, based on new observations and digital reconstruction of the internal anatomy. The result adds one new autapomorphy to the diagnosis of the taxon: ectopterygoid with large boss on the ventral surface and a posteriorly directed process. Additionally new features are recorded: presence of stapes; pterygoid overlapping the vomer, medial sulcus on the dorsal surface of the vomer, parasphenoid–basisphenoid complex covering the ventral surface of the body of the basioccipital. The presence of stapes in Tuarangisaurus keyesi is surprising as they were previously considered absent among elasmosaurids. Phylogenetic analysis indicates a clear elasmosaurid affinity, although there is low resolution within the Elasmosauridae. The previously proposed close relationship with the early Aptian Callawayasaurus colombiensis is rejected.  相似文献   

10.
Mongolian Paleontological Center (MPC) 100/1305 is one of the most complete ankylosaurid skeletons ever collected, and includes much of the postcranial skeleton and numerous in situ osteoderms. This specimen has been referred to Saichania chulsanensis, based on the similarity of the skull compared to the holotype of Saichania, MPC 100/151. However, MPC 100/1305 does not include a skull, and so referral of MPC 100/1305 to Saichania must be based on postcranial characters. Comparison of the postcrania of MPC 100/1305 and MPC 100/151 reveals several differences in the scapula, humerus, and metacarpals, indicating that MPC 100/1305 should not be referred to Saichania. Additionally, although it was previously reported that MPC 100/1305 was collected from the Baruungoyot Formation at Khulsan in Mongolia, collection records indicate that this specimen was instead collected from the Djadokhta Formation at Zamyn Khond. Two ankylosaurid species are known from the Djadokhta Formation of Mongolia and China, Pinacosaurus grangeri and P. mephistocephalus. There are no diagnostic characters in MPC 100/1305 that can be used to refer this specimen to P. grangeri or P. mephistocephalus, and there are a few differences between MPC 100/1305 and Pinacosaurus, such as the number of caudal vertebrae, and morphology of the coracoid, which have uncertain taxonomic implications. At present, MPC 100/1305 is best referred to Ankylosauridae indet., or cf. Pinacosaurus, based on its generally congruent morphology with Pinacosaurus and its provenance from the Djadokhta Formation, in which Pinacosaurus is the only recognized ankylosaurid taxon.  相似文献   

11.
Abstract: A skull and a series of associated cervical vertebrae (ZLJ0112) discovered from the Lower Lufeng Formation (Lower Jurassic) are determined as a juvenile specimen of Lufengosaurus huenei Young 1941 based on amended autapomorphies. Differences between ZLJ0112 and the holotype (sub-adult specimen) are considered as ontogenetic characteristic changes of L. huenei. Since some of these differences are present in other prosauropod dinosaurs (i.e., forms of the maxillary vascular foramen are irregular; the frontal contribution to the dorsal margin of the orbit is substantial; the frontal contribution to the supratemporal fossa is absent; the supratemporal fenestra is visible in lateral view; the supraoccipital inclined at 75 degrees; the parasphenoid rostrum lies level with the occipital condyle; the retroarticular process is short; the axial postzygapophysis project caudally beyond the end of the centrum) they may be common ontogenetic changes in prosauropod dinosaurs.  相似文献   

12.
The fluvio-lacustrine deposits of the ?Cenomanian-Turonian Candeleros Formation in the northwestern fringe of Patagonia have yielded numerous remains of vertebrates, including anurans. A new, partially articulated immature specimen of a small pipimorph pipoid from the upper part of this unit is described herein. Although incompletely preserved, the morphology and proportions of the skeleton are strongly reminiscent of corresponding elements of the holotype of Avitabatrachus uliana from the lower section of the same formation at a nearby locality, thus suggesting that these specimens might represent the same, or a closely related, taxon. The new specimen is a metamorphosing individual: it has a zygapophyseal articulation between the sacrum and the neural arch of a postsacral vertebra that bears conspicuous transverse processes, and four pairs of ribs. These traits are not present in the available material of A. uliana, but they might have occurred within the normal range of variation of a single species. Although the generic and specific allocation of the new material is uncertain, a parsimony analysis performed to clarify the relationships of A. uliana supports a phylogenetic placement of this taxon outside the crown Pipidae. The evidence provided by the Candeleros anurans confirms the presence of stem pipids in Patagonia in the early Late Cretaceous.  相似文献   

13.
This paper describes a new ichthyosaur specimen from the Stonebarrow Marls Member (Charmouth Mudstone Formation) of Charmouth, Dorset. It is substantially complete and various characters of the skull (snout ratios and basioccipital morphology) and appendicular skeleton (the number of digits on the hind paddle, which can be used to infer the number of digits on the front paddle) indicate that the specimen is referable to Ichthyosaurus communis. This provides the first direct evidence for the presence of I. communis in the Pliensbachian, extending the stratigraphic range of the species. The specimen also preserves a probable gullet content, which demonstrates that fish formed part of the diet of this individual.  相似文献   

14.
The holotype of ‘Procheneosaurusconvincens, a juvenile lambeosaurine, is the most complete hadrosaurid known from Kazakhstan. North American species of Procheneosaurus are considered juveniles of Corythosaurus, Hypacrosaurus, and Lambeosaurus, rendering the generic name unusable. A replacement name, Kazaklambia convincens comb. nov., is herein proposed as this specimen is morphologically distinct from other Eurasian taxa and known juvenile lambeosaurines at a similar ontogenetic stage in having a prefrontal process of the postorbital with a dorsal thickening forming a dome lateral to the frontal dome, doming of the nasal anterodorsal to the orbit, and a frontal length/width ratio <1. The juvenile status of Kazaklambia makes phylogenetic placement difficult; however, morphometric and morphological information (particularly in relation to the hollow cranial crest and the length of the frontal) suggest a close affiliation with the basal lambeosaurines Amurosaurus and Tsintaosaurus, and support the hypothesis for an Asian origin for Lambeosaurinae.  相似文献   

15.
A New Titanosaurian Sauropod from Late Cretaceous of Nei Mongol, China   总被引:2,自引:0,他引:2  
A specimen collected from the Upper Cretaceous Erlian Formation of Nei Mongol (Inner Mongolia), China, represents a new genus and species of titanosaurian sauropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises several dorsal, sacral, and caudal vertebrae, several dorsal ribs, one anterior chevron, and much of the pelvis. Diagnostic features of the new species include a thick ridge extending down from the postzygapophysis on the lateral surface of the neural arch of the posterior dorsal vertebrae, a transversely oriented accessory lamina present between the anterior centroparapophyseal limina and the lateral centraprezygapophyseal lamina of the posterior dorsal vertebrae, long, anteroventrally directed caudal rib that bears two distinctive fossae on its posterior margin on the anterior caudal vertebrae and a prominent vertical ridge above the pubic peduncle on the medial surface of the ilium, among others. The ilium is pneumatic, a feature not common among non-avian dinosaurs. The new taxon has an unusual combination of primitive and derived character states. Preliminary character analysis shows a complex character distribution within the Titanosauriformes. Recent titanosauriform discoveries suggest that a significant radiation occurred in Asia early in the titanosauriform evolution.  相似文献   

16.
A new species of the sail-crested pterosaur Tupuxuara is described from the Santana Formation of Brazil, Tupuxuara deliradamus sp. nov. The holotype, a partial skull, and a larger, partial skull referred to the same taxon differs from Tupuxuara leonardii by having a nasoantorbital fenestra with an acutely-angled posterior border with a long, straight posterodorsal margin, a reclined cranium, and an orbit situated entirely in the ventral half of the nasoantorbitral fenestra. Unfortunately, neither specimen is comparable with the fragmentary rostrum representing Tupuxuara longicristatus. In addition, resolution of a recent nomenclatural problem over the correct name for the clade containing Tupuxuara and its sister taxon, Thalassodromeus, is provided. Both genera are used by different authors as the nomenclatural basis for the group, but “Tupuxuaridae” has never been explicitly erected as a new taxon, and therefore fails to meet ICZN criteria that new taxa are only valid if authors clearly indicate their intention to establish new names. By contrast, “Thalassodrominae” was explicitly erected as a name for the Thalassodromeus + Tupuxuara clade, thereby fulfilling all ICZN requirements for naming of a new taxon and making Thalassodromeus stand as the type genus for this group.  相似文献   

17.
The Upper Cretaceous of Africa has produced a diverse fauna of mosasaurs, including the highly specialized, long-jawed Pluridens. The type of Pluridens walkeri comes from the Maastrichtian Farin-Doutchi Formation of Niger, with a second, referred specimen coming from the Campanian section of the Campanian-Maastrichtian Nkporo Shale near Calabar, in southern Nigeria. Comparisons of this referred specimen with the holotype suggest that it represents a distinct and more primitive species. The Calabar jaw resembles P. walkeri in being long and narrow anteriorly with a shallow subdental shelf, and in having small, numerous, recurved teeth with medially positioned replacement pits. However, it lacks many of the derived features that characterize Pluridens walkeri, such as the extremely long and straight jaw, the extreme lateral protrusion and subcircular section of the dentary, strong transverse expansion of the dental thecae, and extreme reduction and increase in number of the teeth. The Calabar Pluridens is therefore referred to a new species, Pluridens calabaria. Following recent studies, Pluridens is considered to represent a highly derived member of the Halisaurinae. The marked differences between the Campanian and Maastrichtian species of the genus underscore the rapid pace of mosasaur evolution during the Cretaceous.  相似文献   

18.
A fragmentary ichthyosaur specimen collected in situ at Castle Top Quarry in Nettleton, Lincolnshire, UK from exposures of the Lower Cretaceous (Berriasian) Spilsby Sandstone Formation (Subcraspedites ?preplicomphalus Zone) is reported. In general, Early Cretaceous ichthyosaurs from the Berriasian to Barremian are poorly understood. Despite the fragmentary nature of the described specimen, it is the first ichthyosaur reported from this specific zone and adds to the literature another rare ichthyosaur from the Berriasian.  相似文献   

19.
The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments.  相似文献   

20.
Upper Cretaceous vertebrate accumulations from the Adamantina Formation are known due to their high taxonomic diversity. On the other hand, taphonomic analyses still are rare, limiting the understanding of processes related to the biostratinomic and fossildiagenetic histories of this lithostratigraphic unit. In 2005, fossils were collected from an outcrop located at Jales municipality, state of São Paulo, southeastern Brazil. From this outcrop, a well-preserved Baurusuchus was recovered, which displays a peculiar set of taphonomic signatures. This paper identifies and interprets taphonomic features of a specimen of Baurusuchus (Crocodyliformes, Baurusuchidae; UFRJ DG 418-R) from the Adamantina Formation (Upper Cretaceous of the Bauru Basin), in Jales municipality, state of São Paulo. Brief taphonomic comparisons with other specimens previously studied (crocodiles and dinosaurs) and a lithofaciological analysis of the outcrop were undertaken in order to enhance the knowledge of the stratigraphy and paleoenvironment and improve the time resolution for the Adamantina Formation in the region of Jales. Furthermore, paleoecological data were interpreted based on the taphonomic analysis.The fossil is composed of an articulated segment of vertebral column, ribs, part of the pelvic girdle and gastralia. There is no hydraulic equivalence between both the specimen of Baurusuchus and the size of quartz grain predominant in the fossiliferous layer, suggesting death in situ or short transport as a “water carcass”. Teeth marks identified on the pubes were assigned to a small/juvenile baurusuchid crocodyliform or a theropod dinosaur. The repositioning of some elements (ribs and dorsal osteoderms) is suggestive of mummification. Desiccation marks were observed and attributed to the stage 1 of weathering. These features suggest subaerial exposure of the carcass prior to burial, however, probably after the mummification. On the other hand, the subaerial exposure was short, because the individual was not fully disarticulated. Furthermore, the degrees of articulation and preservation of the specimen nullify the hypothesis of reworking.Lithofaciological and taphonomic analyses suggest a well-drained floodplain as the burial environment and an arid or semi-arid climate in the region of Jales outcrop. In addition, the taphonomic signatures seem to indicate a time resolution about 100–104 years for the layer where the crocodyliforms were collected, but it seems to have, within this time limit, temporal-mixing among terrestrial crocodiles and dinosaurs collected from the same layer, suggesting time-averaging in this outcrop.  相似文献   

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