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1.
The origin of the genus Bos is a debated issue. From ∼ 0.5 Ma until historic times, the genus is well known in the Eurasian large mammal assemblages, where it is represented by Bos primigenius. This species has a highly derived cranial anatomy that shows important morphological differences from other Plio-Pleistocene Eurasian genera of the tribe Bovini such as Leptobos, Bison, Proamphibos-Hemibos, and Bubalus. The oldest clear evidence of Bos is the skull fragment ASB-198-1 from the middle Pleistocene (∼ 0.6-0.8 Ma) site of Asbole (Lower Awash Valley, Ethiopia). The first appearance of Bos in Europe is at the site of Venosa-Notarchirico, Italy (∼ 0.5-0.6 Ma). Although the origin of Bos has traditionally been connected with Leptobos and Bison, after a detailed anatomical and morphometric study we propose here a different origin, connecting the middle Pleistocene Eurasian forms of B. primigenius with the African Late Pliocene and early Pleistocene large size member of the tribe Bovini Pelorovis sensu stricto. The dispersal of the Bos lineage in Western Europe during middle Pleistocene times seems to coincide with the arrival of the Acheulean tool technology in this continent.  相似文献   

2.
Fossil skeletal remains of the Afro–Asian labeonine genus Garra are described on the basis of more than 300 specimens from the Pliocene diatomites of Jradzor, Central Armenia. Extant species of the genus Garra exhibit a wide distribution ranging from China, South Asia, and Borneo, through Middle East, the Arabian Peninsula, and Africa, where these fishes inhabit a variety of freshwater biotopes. The Pliocene fossils from the Jradzor site provide the first evidence of Garra in the fossil record. The inadequate preservation of the available material did not allow a detailed taxonomic attribution at the species level. Remains of Garra sp. were found associated to a moderately diverse assemblage, which includes also remains of additional freshwater fishes (Capoeta sp., Leuciscus cf. souffia, Leuciscus sp.), amphibians (Pelophylax cf. ridibundus), and large mammals (Rhinocerotidae indet.), as well as of monocotyledon plants. According to the diatom flora, the diatomites of Jradzor can be assigned to the Pliocene and were deposited in a freshwater lacustrine setting that was characterized by high productivity, eutrophic conditions, and standing macrophyte vegetation along the littoral zone. The presence of abundant resorptive pharyngeal teeth suggests that the Pliocene palaeolake of Jradzor was characterized by resident populations of Garra. Extant species of this genus are currently absent in the freshwaters of Armenia. The record of the genus Garra from Jradzor suggests that the Araks–Kura River drainage was connected, at least in part, with the Euphrates and Tigris River drainage during the Pliocene. The extinction of Garra (and other thermophilous taxa) from Araks and Kura River drainage was probably due to Plio-Pleistocene tectonic uplift of the Armenian Highland and consequent progressive climate cooling.  相似文献   

3.
The marine Pliocene-Pleistocene sequence of Iceland contains four main assemblages of mollusks: (1) Venerupis rhomboides et al., (2) Serripes groenlandicus et al., (3) Portlandia arctica et al., and (4) recent species. Analysis of the stratigraphic distribution of 100 species shows that the percentage of extinct species in the assemblages decreases from 40% in the oldest to 4–8% in the youngest. There is also a change from south boreal affinities in the oldest assemblage to north boreal in the youngest. The molluskan assemblages of Iceland can be correlated with Pliocene-Pleistocene assemblages of England and North Europe: (1) Coralline Crag, (2) Red Crag—lower part of the Icenian strata, (3) upper part of the Icenian-Cromerian strata. The upper boundaries of the Pliocene and Eopleistocene are defined by paleomagnetic data and radiological dating. The upper boundary of the Pliocene presumably should be established in the basalts lying between the Tjörnes and the Breidavik deposits by changes between the second and third mollusk assemblages and by the Gilsa paleomagnetic event.  相似文献   

4.
Remains of the cercopithecid Mesopithecus monspessulanus are relatively rare. Two previously unpublished mandibles of M. monspessulanus (housed in the Basel Naturhistorisches Museum), from the Italian locality of Villafranca d’Asti are described. These remains belong to the assemblage of the Triversa Faunal Unit, dated to the Early Villafranchian, that is to the unit MN16a (Middle Pliocene) of the European mammal biochronology. According to this recently revised biochronological attribution, Villafranca d’Asti records the last Mesopithecus occurrence in Europe. The NOW (Neogene Old World) database has been used as a basis to evaluate both the mammalian faunal and the palaeoenvironmental context at the time surrounding this last occurrence of Mesopithecus. The comparison (taxonomical composition and ungulate hypsodonty), between the Villafranca d’Asti assemblage and other Plio-Pleistocene mammal communities of Europe, shows that the extinction of Mesopithecus is related to a faunal turnover and a change toward more open landscapes during the Early–Middle Villafranchian transition. This is consistent with the “Elephant-Equus event”, that occurred in Europe at 2.5 Ma. Furthermore, the co-occurring but more terrestrial cercopithecid Macaca crossed this faunal turnover. This strengthens the assumption that the latest Mesopithecus species, M. monspessulanus, had woodland-oriented adaptations. Editorial handling: J.-P. Billon-Bruyat & M. Chiari (Guest)  相似文献   

5.
《Quaternary Science Reviews》2005,24(10-11):1287-1301
Human occupants of Europe shared food resources with a number of larger Carnivora, and their coexistence with two lion-sized felids, the lion and the scimitar-toothed machairodont Homotherium latidens, poses intriguing paleoecological problems. We investigate the ecology of Homotherium latidens using an exceptional sample of postcrania from the Spanish Early Pleistocene site of Incarcal, making comparisons with modern cats and with other machairodont species. Evidence of cursorial adaptations in Homotherium suggests a hunting technique different from modern cats or smilodontine sabre-tooths. Some, like reduction of the claws, would have limited the ability of individual homotheres to bring down large prey, implying group action. Homotherium would also have been disadvantaged in direct confrontation with Pleistocene lions by smaller body mass, reduced forepaw muscle strength, smaller claws and more fragile dentition. Its hunting technique would have worked best in more open habitats, but competition from lions would have forced it to seek moderate cover. Among factors that could de-stabilise coexistence of the two big cat species in Pleistocene Europe we invoke a decrease in environmental mosaicism associated with stepped climatic change over the last million years, and the increased importance of humans within the larger predator guild.  相似文献   

6.
Climate change during the Quaternary played an important role in the distribution of extant plants. Herein, cone scales of Cedrus (Pinaceae) were uncovered from the Upper Pliocene Sanying Formation, Longmen Village, Yongping County of Yunnan Province in southwestern China. Detailed comparisons show that these fossils all belong to the genus Cedrus (Pinaceae), and a new species is proposed, Cedrus angusta sp. nov. This find expands the known distribution of Cedrus during the Late Pliocene to Yunnan, where the genus no longer exists in natural forests. Based on the analysis of reconstructed Neogene climate data, we suggest that the intensification of the East Asian winter monsoon during the Quaternary may have dramatically increased seasonality and given rise to a much drier winter in Yunnan. Combined with information on Cedrus fossil records and its seed physiology, we conclude that the intensification of a drier climate after the Late Pliocene may have prevented the survival of Cedrus seedlings, leading to the eventual disappearance of Cedrus in western Yunnan. This study indicates that the topography in southwestern China acted as a vital refuge for many plants during the Quaternary, but that other species gradually disappeared due to the intensification of the monsoonal climate.  相似文献   

7.
A fossiliferous fissure filling at Les Alleveys, Mormont (Canton de Vaud, Switzerland), sampled in 1986 and representing part of the same fissure system as that excavated by Chavannes and Morlot in 1852, contains an Eocene mammal fauna of 27 species-group taxa, dominated by micromammals. Unlike the 1852 collection, thanks to screenwashing techniques, there is an abundance of rodents, especially theridomyids, which provide new information on the early radiation of the family. The homogeneity of the new assemblage suggests that the Les Alleveys fauna is entirely Bartonian in age and not as previously thought mixed with Priabonian elements. Bartonian faunas rich in a diversity of micromammals are relatively uncommon in Europe outside the Quercy region of France and this is the first such fauna from Mormont and from Switzerland. The rodents are here treated systematically and a new species of Elfomys, E. engesseri sp. nov. is described. A few teeth of an undescribed species tentatively referred to ‘Protadelomys’ provide an archaic element, whereas a single milk premolar of Patriotheridomys? suggests a considerable range for the genus prior to the middle Priabonian. The Les Alleveys theridomyids, together with a range of other early family members are analysed cladistically. The analysis confirms both the distinctness of the subfamily Remyinae and that the genus Estellomys is a primitive theridomyine; it shows Paradelomys to be a primitive member of the Columbomyinae, but finds no dental synapomorphies to support the Issiodoromyinae as represented by Elfomys and Pseudoltinomys.  相似文献   

8.
《Gondwana Research》2014,25(3):1186-1201
The modern Koala Phascolarctos cinereus is the last surviving member of a once diverse family Phascolarctidae (Marsupialia, Phascolarctomorphia). Nine genera and at least 16 species of koala are known. Late Oligocene sediments of central Australia record the oldest fossils and highest species diversity. Five species are known from the early to middle Miocene rainforest assemblages of the Riversleigh World Heritage Area, Queensland. With the onset of dryer conditions after the middle Miocene climatic optimum (~ 16 Ma), rainforest habitats contracted resulting in the apparent extinction of three koala lineages (Litokoala, Nimiokoala, Priscakoala). Phascolarctos first appears in the fossil record during the Pliocene and the modern species around 350 ka. Despite a dramatic decline in taxonomic diversity to a single extant species, the fossil record indicates that at most only three koala species coexisted in any given faunal assemblage throughout their 24 million year history. Within these assemblages, the vast majority of extinct koalas are extremely rare (some known from only a single specimen) which may reflect a general rarity within their palaeohabitats compared with the modern species which is represented by an estimated 400,000 individuals spread over most of eastern mainland Australia. Be that as it may, P. cinereus, although once geographically more widespread, occurring for example in Western Australia in the Pleistocene, underwent significant range contractions and localized population extinctions during the stressful climatic conditions of the late Pleistocene and more recently through human-induced habitat destruction. Combined with threats of disease, reduced genetic diversity and climate change, the survival of this iconic Australian marsupial is arguably a cause for concern.  相似文献   

9.
The fossil hamster (an isolated m1) from the early Early Pleistocene Houhecun fauna found in Dali, Weinan, Shaanxi, China, originally identified as Kowalskia dalinica, is reexamined. Its unique characters, including a high crown, very small degree of division of the anteroconid and relatively large size, is taken as evidence of its attribution to the genus Sinocricetus, instead of Kowalskia (junior synonym of Neocricetodon) suggested by the original author. Therefore, the specific name of the species established based on this m1 should be changed to S. dalinicus (Wang, 1988). S. dalinicus probably has a relatively close affinity with S. major Li, 2010, but there are still some obvious differences of characters between them. The discovery of S. dalinicus in the Houhecun fauna confirms that the genus indeed survived into the Pleistocene. A broken hamster mandible, which was found in Weinan, Shaanxi, China and belongs to the late Pliocene Youhe fauna, is also described here and identified as Tscherskia sp.. This specimen represents the earliest Tscherskia in Asia so far, but fails to refute the hypothesis that the genus originated in Europe during the early Pliocene.  相似文献   

10.
Although the European dinosaur succession during the latest Cretaceous and its relationship with the Cretaceous-Paleogene (K-Pg) mass extinction has been the focus of recent work, other continental vertebrates, such as crocodylomorphs, have received less attention. The European continental record of crocodylomorphs in general, and of eusuchians in particular, is relatively dense through the Maastrichtian until the K-Pg boundary. Traditionally it has been argued that continental crocodylomorphs were minimally impacted by the K-Pg extinction, but they were substantially affected in Europe with the disappearance of endemic eusuchians such as Hylaeochampsidae, Allodaposuchus and their close relatives, and non-eusuchians such as Doratodon or Theriosuchus. Despite extensive sampling in Danian continental deposits, only scarce fragmentary crocodylomorph remains have been cited. It is not until the late Paleocene and Eocene that a recovery in continental crocodylomorphs is observed. The presence of taxa such as planocraniids, the alligatoroids Diplocynodon and Hassiacosuchus, and stem crocodyloids during this period provide the first reliable continental records of Crocodylia in Europe and is best explained by post-extinction immigration from Asia or North America. By contrast, marine forms such as Thoracosaurus are found on both sides of the K-Pg boundary in Europe. The adaptation of these marine animals to different environments, from shallow seas to more transitional or fluvial environments, could be the key to their success and survival across the K-Pg boundary, as seen in other marine crocodylomorph clades such as Dyrosauridae.  相似文献   

11.
The aim of this paper is to reconstruct the variation in the vegetation during the Holocene on the Northern Meseta through pollen analysis in order to determine the roles played by Pinus and Quercus in the vegetation dynamics. A new pollen sequence, Camporredondo, in the centre of the Duero River basin, Northern Meseta, Spain, is presented. It covers the period from ca 11,250 to 1630 cal yr BP. The Camporredondo sequence suggests that a forested landscape existed from the start of the Holocene with values of approximately 90% of arboreal pollen mainly derived from pine forests in the surroundings. Shortly after 8600 cal yr BP pine forests continued to dominate but the landscape became more open while stands of deciduous and evergreen Quercus developed. Around 4500 years ago, coinciding with the start of a period of less rainfall, the lacustrine areas in these territories began to silt up, favouring hydrophytic taxa such as Salix or Cyperaceae. A comparison with other Holocene pollen sequences from the central-eastern part of the Northern Meseta confirms the presence of pine forests in the whole region since the early Holocene. We discuss the variations of the general vegetation pattern in terms of the local or regional conditions at each site. Throughout the Holocene, Pinus forests dominated at least in the eastern half of the Duero River basin. The typical Holocene substitution of conifers by broadleaf species as recorded in other parts of the Iberian Peninsula and Europe, never takes place in the study area. The results of the present study suggest that the current view of the potential vegetation in the Spanish Northern Meseta should be reviewed.  相似文献   

12.
A representative early Bajocian belemnite assemblage containing four genera and 11 species (two identified in open nomenclature and five new: Eocylindroteuthis weisi sp. nov., E. mariottii sp. nov., E. yarkovi sp. nov., Homaloteuthis volgogradensis sp. nov., and Hastites orphana sp. nov.) is described from a section near the Dubovoi hamlet in the Greater Don Bend area (southern termination of the Don-Medveditsa dislocations). Some members of this assemblage were previously known from the Caucasus; however, it is possible to state the discovery of a Euroboreal belemnite fauna completely new for Russia, previously described only from Central Europe and almost unknown for Eastern Europe. The age of the studied assemblage collected from the lower part of the section, traditionally considered to be Upper Bajocian, corresponds to the Laeviuscula ammonite Chronozone and is the first reliable evidence of marine settings in the Volga Region for the early Bajocian. Two new biostratigraphic units, Beds with Eocylindroteuthis weisi and Beds with H. orphana, are introduced. The former, judging from the occurrences of its characteristic species in Western Europe, has a broad correlative potential. On the basis of datings obtained, the scheme of the formational subdivision of the Middle Jurassic of the Volga Region near Volgograd is revised, and the formerly abandoned Bakhtemir Formation is reinstated as a valid unit. Its total range is reestablished as corresponding to the upper part of the Discites (?)/Laeviuscula Chronozone (lower Bajocian)–Garantiana Chronozone (upper Bajocian).  相似文献   

13.
Marine Pliocene and Pleistocene beds of the Boso Peninsula are more than 3 km thick and contain abundant fauna and flora. Almost continuous deposition occurred at a high sedimentation rate from the early part of the last reversed geomagnetic polarity epoch to the middle part of the last normal polarity epoch. Several microbiostratigraphic marker horizons enable correlation with deep-sea sediments fully independent of magnetostratigraphic zonation. Globorotalia truncatulinoides first occurs below the normal polarity event BO-B-2, which is correlated with the Olduvai Event in deep-sea sediments, whereas the disappearance of discoasters and the first occurrence of Gephyrocapsa caribbeanica are recognized at the top of BO-B-2, and the first occurrence of Gephyrocapsa oceanica is recognized at the top of BO-B-1. Based on the correlation of the Boso section with Mediterranean stage stratotype sections, the transition from the Pliocene to Pleistocene is estimated to be in the lower part of the Kazusa Group.  相似文献   

14.
Early and early middle Cenomanian turrilitid ammonoids from the upper part of the Albian–Cenomanian Aitamir Formation of the Koppeh Dagh (northeast Iran) are described, illustrated and placed in an integrated stratigraphic context. The Aitamir Formation represents a graded siliciclastic shelf system and the turrilitid faunas comprise eight different species. Representatives of Mariella and Hypoturrilites have been recorded from a lower Cenomanian horizon in the Ghorghoreh section, corresponding to the Mantelliceras mantelli ammonite biozone. Mariella bicarinata (Kner, 1852) and H. wiedmanni Collignon, 1964 are recorded for the first time from Iran and the larger palaeobiogeographical area, respectively. The horizons with Turrilites costatus and T. scheuchzerianus at Taherabad can be dated as early middle Cenomanian Acanthoceras rhotomagense Zone, T. costatus Subzone. Concentrations of ammonoids commonly occur above regressive sandstone units, at the transition into overlying deeper marine shale units and in their lower parts, respectively. Such intervals represent the early transgressive systems tract of depositional sequences and may thus be regarded as early transgressive, slightly condensed shell accumulations. The uppermost lower to lower middle Cenomanian succession at Taherabad is of striking bio-, event- and sequence stratigraphic similarity to contemporaneous sections in northwest Europe. It reflects deposition during falling and low sea-level associated with the latest early Cenomanian unconformity SB Ce 3 as well as the pulsating transgressive development during the early middle Cenomanian, including levels correlative to the arlesiensis, primus and Mid-Cenomanian events in northwest Europe.  相似文献   

15.
The three sedimentary units infilling Lake George provide the longest quasi-continuous sedimentary record of any Australian lake basin. A combination of cosmogenic nuclide burial, magnetostratigraphy and biostratigraphic dating techniques previously has shown that the basal (fluvial) unit, the Gearys Gap Formation, began accumulating at ca 4 Ma, in the early Pliocene (Zanclean), and (ii) deposition had ceased by ca 3 Ma, in the mid-late Pliocene (Piacenzian). The same techniques confirm the middle unit, the (fluvio-lacustrine) Ondyong Point Formation began accumulating in the late Pliocene and deposition continued into the earliest Pleistocene (Gelasian) when a shallow but probably laterally extensive freshwater lake extended across the drillhole site. Our data provide a minimum Gelasian age for tectonic blockage of former spillway(s) and formation of paleo-Lake George. Whether this was the earliest lake to form within the basin is unknown, since the dated intervals are separated by a ferric hardpan, interpreted as representing a prolonged period of erosion or non-deposition. Temperate rainforest angiosperms including Nothofagus growing during the late Pliocene had been extirpated or become extinct during this interval, although a number of gymnosperms, now endemic to New Caledonia, New Guinea, New Zealand and Tasmania still survived in the otherwise sclerophyll-dominated vegetation. The succession of plant communities is considered to be due to effectively drier local conditions, which in turn reflect regional aridification during the Plio-Pleistocene transition, despite the formation of a freshwater lake across the basin. The sequence provides a reliable framework for recognising and correlating Plio-Pleistocene deposits elsewhere on the Southern Highlands.  相似文献   

16.
Knowledge of the main aspects of the Great American Biotic Interchange (GABI) concerning the glyptodontine Glyptodontidae (Xenarthra) is very scarce. A bidirectional dispersal process was recently proposed for this clade, with the presence of the North American genus Glyptotherium Osborn recognized in latest Pleistocene sediments of northern South America (Venezuela and Brazil). However, the earliest stages of this paleobiogeographical process remain poorly understood, mainly because of the limited fossil record on this clade in late Pliocene sediments. The goals of this contribution are: a) to present and describe the first record of a glyptodontine glyptodontid from the late Pliocene of northern South America, tentatively assigned to a new species of Boreostemma Carlini et al. (Boreostemma? sp. nov); and b) to analyze its paleobiogeographical implications with respect to the GABI. This new material was recovered from the San Gregorio Formation (late Pliocene, prior the GABI) in northern Venezuela, where it is represented by several osteoderms of the dorsal carapace. A comparison among the three known late Pliocene glyptodontine glyptodontids of a) southern South America (Paraglyptodon), b) northern South America (Boreostemma), and c) southern North America ("Glyptotherium"), reveals a series of shared characters between (b) and (c), not present in (a). The most important of these shared characters in (b) and (c) are: all the osteoderms present a great development of the central figure, which is always larger than the peripherals; the sulcus that delimits the central and peripheral figures is narrower and shallower; and all the osteoderms present are relatively thin. This evidence suggests that the lineage of Glyptodontinae which participated in the GABI and subsequently diversified in North America originated in northern South America. Moreover, the evident morphological differences between these glyptodontines with respect to the southern South American forms show a significant separation of both lineages since at least latest Miocene-early Pliocene.  相似文献   

17.
The Paleocene-Early Eocene larger benthic foraminifera (LBF) in the far eastern Neo-Tethyan Ocean of Tibet still remain poorly known. Here, we present a novel, high-resolution larger foraminiferal biozonation from the shallow-water limestones in Tibet, which will improve our current understanding of the larger foraminiferal evolution in the eastern Neo-Tethyan Ocean. Based on one continuous section at Tingri and three separate sections at Gamba, ten Shallow Benthic Zones (SBZ 1–10) have been designated in Tibet by following the principle of Oppel Zone. In contrast to those in Europe, the Paleocene LBFs in Tibet are characterized by high diversification of Lockhartia, Kathina, Daviesina, Miscellanea, Ranikothalia, and Operculina and show progressively increasing diversity of genera and species during SBZ 2–5. Adult dimorphism and large shell size of some LBFs as well as differentiation of the diversity between genera and species initiated as early as SBZ 3. It suggests that the occurrence of the Larger Foraminifera Turnover (LFT) was probably not synchronous in the entire Neo-Tethyan Ocean, because in Europe, the LFT was generally thought to occur at the beginning of SBZ 5. During the Early Eocene, the LBFs in Tibet decreased markedly on the generic level and increased on the species level, and some new genera (Alveolina, Orbitolites, Nummulites, Assilina, Discocyclina) have gained predominance in Tibet. It is nearly identical to the evolution of the LBFs in Europe and indicates a high-degree homogenization of the LBFs in the entire Neo-Tethyan Ocean. Furthermore, the Paleocene-Eocene (P-E) boundary in shallow-water environments has been clearly identified by us, and it is situated in the upper part of SBZ 5 and associated with no evident biotic turnover of shallow benthic foraminiferal communities. The possible diachroneity of the LFT in the Neo-Tethyan Ocean and the evident lagging of the Paleocene-Eocene Thermal Maximum (PETM) behind the LFT imply that the LFT could only be the result of a natural evolutionary process and has no linkage with the PETM. Notably, a transient but distinct larger foraminiferal extinction and origination (LFEO) event has been found in Tibet, which is characterized by a sudden disappearance of all Paleocene lamellar-perforate LBFs, such as Lockhartia, Kathina, Daviesina, Miscellanea, Ranikothalia, and Operculina, and the initial dominance of the Early Eocene porcellaneous-walled Alveolina. The LFEO marks the boundary between SBZ 5 and 6, and might only occur in the low-latitude areas of the Neo-Tethyan Ocean. Surprisingly, the LFEO coincides with the initial recovery of the Carbon Isotope Excursion (CIE), and their synchronicity implies that some possible mechanisms causing the rapid recovery of the CIE probably had also led to the LFEO in the shallow-water environments.  相似文献   

18.
West Anatolia, together with the Aegean Sea and the easternmost part of Europe, is one of the best examples of continental extensional tectonics. It is a complex area bounded by the Aegean–Cyprus Arc to the south and the North Anatolian Fault Zone (NAFZ) to the north. Within this complex and enigmatic framework, the Sandıklı Graben (10 km wide, 30 km long) has formed at the eastern continuation of the Western Anatolian extensional province at the north‐northwestward edge of the Isparta Angle. Recent studies have suggested that the horst–graben structures in West Anatolia formed in two distinct extensional phases. According to this model the first phase of extension commenced in the Early–Middle Miocene and the last, which is accepted as the onset of neotectonic regime, in Early Pliocene. However, it is controversial whether two‐phase extension was separated by a short period of erosion or compression during Late Miocene–Early Pliocene. Both field observations and kinematic analysis imply that the Sandıklı Graben has existed since the Late Pliocene, with biaxial extension on its margins which does not necessarily indicate rotation of regional stress distribution in time. Although the graben formed later in the neotectonic period, the commencement of extension in the area could be Early Pliocene (c. 5 Ma) following a severe but short time of erosion at the end of Late Miocene. The onset of the extensional regime might be due to the initiation of westward motion of Anatolian Platelet along the NAFZ that could be triggered by the higher rate of subduction at the east Aegean–Cyprus Arc in the south of the Aegean Sea. Copyright © 2003 John Wiley & Sons, Ltd.  相似文献   

19.
The Sierra Nevada of southern Spain is a landscape with a rich biological and cultural heritage. The range was extensively glaciated during the late Pleistocene. However, the postglacial paleoecologic history of the highest range in southern Europe is nearly completely unknown. Here we use sediments from a small lake above present treeline – Laguna de Río Seco at 3020 m elevation – in a paleoecological study documenting over 11,500 calendar years of vegetation, fire and climate change, addressing ecological and paleoclimatic issues unique to this area through comparison with regional paleoecological sequences. The early record is dominated by Pinus pollen, with Betula, deciduous Quercus, and grasses, with an understory of shrubs. It is unlikely that pine trees grew around the lake, and fire was relatively unimportant at this site during this period. Aquatic microfossils indicate that the wettest conditions and highest lake levels at Laguna de Río Seco occurred before 7800 cal yr BP. This is in contrast to lower elevation sites, where wettest conditions occurred after ca 7800. Greater differences in early Holocene seasonal insolation may have translated to greater snowpack and subsequently higher lake levels at higher elevations, but not necessarily at lower elevations, where higher evaporation rates prevailed. With declining seasonality after ca 8000 cal yr BP, but continuing summer precipitation, lake levels at the highest elevation site remained high, but lake levels at lower elevation sites increased as evaporation rates declined. Drier conditions commenced regionally after ca 5700 cal yr BP, shown at Laguna de Río Seco by declines in wetland pollen, and increases in high elevation steppe shrubs common today (Juniperus, Artemisia, and others). The disappearance or decline of mesophytes, such as Betula from ca 4000 cal yr BP is part of a regional depletion in Mediterranean Spain and elsewhere in Europe from the mid to late Holocene. On the other hand, Castanea sativa increased in Laguna de Río Seco record after ca 4000 cal yr BP, and especially in post-Roman times, probably due to arboriculture. Though not as important at high than at low elevations, fire occurrence was elevated, particularly after ca 3700 years ago, in response to regional human population expansion. The local and regional impact of humans increased substantially after ca 2700 years ago, with the loss of Pinus forest within the mountain range, increases in evidence of pasturing herbivores around the lake, and Olea cultivation at lower elevations. Though human impact was not as extensive at high elevation as at lower elevation sites in southern Iberia, this record confirms that even remote sites were not free of direct human influence during the Holocene.  相似文献   

20.
We provide a synopsis of ~ 60 million years of life history in Neotropical lowlands, based on a comprehensive survey of the Cenozoic deposits along the Quebrada Cachiyacu near Contamana in Peruvian Amazonia. The 34 fossil-bearing localities identified have yielded a diversity of fossil remains, including vertebrates, mollusks, arthropods, plant fossils, and microorganisms, ranging from the early Paleocene to the late Miocene–?Pliocene (> 20 successive levels). This Cenozoic series includes the base of the Huchpayacu Formation (Fm.; early Paleocene; lacustrine/fluvial environments; charophyte-dominated assemblage), the Pozo Fm. (middle + ?late Eocene; marine then freshwater environments; most diversified biomes), and complete sections for the Chambira Fm. (late Oligocene–late early Miocene; freshwater environments; vertebrate-dominated faunas), the Pebas Fm. (late early to early late Miocene; freshwater environments with an increasing marine influence; excellent fossil record), and Ipururo Fm. (late Miocene–?Pliocene; fully fluvial environments; virtually no fossils preserved). At least 485 fossil species are recognized in the Contamana area (~ 250 ‘plants’, ~ 212 animals, and 23 foraminifera). Based on taxonomic lists from each stratigraphic interval, high-level taxonomic diversity remained fairly constant throughout the middle Eocene–Miocene interval (8-12 classes), ordinal diversity fluctuated to a greater degree, and family/species diversity generally declined, with a drastic drop in the early Miocene. The Paleocene–?Pliocene fossil assemblages from Contamana attest at least to four biogeographic histories inherited from (i) Mesozoic Gondwanan times, (ii) the Panamerican realm prior to (iii) the time of South America’s Cenozoic “splendid isolation”, and (iv) Neotropical ecosystems in the Americas. No direct evidence of any North American terrestrial immigrant has yet been recognized in the Miocene record at Contamana.  相似文献   

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