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1.
Well-preserved skulls of Late Pleistocene Panthera spelaea are described from two geographically distant regions of Russia situated in the Russian Plain and Northern Siberia. The resemblance of the studied material with contemporaneous fossil lions from Alaska is established. The cranial morphology of P. spelaea confirms its phylogenetic position within the lion group, whereas many of the characters supporting the cave lion's relationship with the tiger are primitive. Both living and cave lions are advanced relative to the Middle Pleistocene P. fossilis in having larger upper incisors and more inflated bullae. P. spelaea also possesses the following advanced characters in common with the living lion: widened muzzle in the canine and P2 areas, wide nasal bones, relatively short pre-orbital part of the skull, laterally widened mastoid area, V-shaped form of the posterior tip of the frontal process of maxillary bone, and reduced cusp on the protocone bulge of P4. These conditions establish P. spelaea as a distinct species.  相似文献   

2.
Upper Pleistocene remains of the Ice Age steppe lion Panthera leo spelaea (Goldfuss, 1810) have been found in the Perick Caves, Sauerland Karst, NW Germany. Bones from many hyenas and their imported prey dating from the Lower to Middle Weichselian have also been recovered from the Perick Cave hyena den. These are commonly cracked or exhibit deep chew marks. The absence of lion cub bones, in contrast to hyena and cave bear cub remains in the Perick Caves, and other caves of northern Germany, excludes the possibility that P. leo spelaea used the cave for raising cubs. Only in the Wilhelms Cave was a single skeleton of a cub found in a hyena den. Evidence of the chewing, nibbling and cracking of lion bones and crania must have resulted from the importation and destruction of lion carcasses (4% of the prey fauna). Similar evidence was preserved at other hyena den caves and open air sites in Germany. The bone material from the Perick and other Central European caves points to antagonistic hyena and lion conflicts, similar to clashes of their modern African relatives.  相似文献   

3.
Bone remains and a trackway of Pantheraichnus bottropensis nov. ichg. ichnsp. of the Late Pleistocene lion Panthera leo spelaea (Goldfuss, 1810) have been recovered from Bottrop and other open air sites in northern Germany. Some of these bones are from open air hyena den sites. A relative high proportion of lion bones (20%) exhibit bite, chew or nibble marks, or bone crushing and nibbling caused by a large carnivore. Repeated patterns of similar bone damage have been compared to bone remains found at hyena dens in gypsum karst areas and cave sites in northern Germany. Ice Age spotted hyenas have been the main antagonists and the main scavengers on lion carcasses. The remains appear to have been imported often by hyenas into their communal dens, supporting the theory of strong hyena-lion antagonism, similar to the well documented antagonism between modern African lions and spotted hyenas. Most of the lion bones from the open air hyena den at Bottrop are probably a result of such antagonism, as are the rare remains of these carnivores found within large hyena prey bone accumulations along the Pleistocene rivers. The Emscher River terrace also has the largest quantity of hyena remains from open air river terrace sites in northern Germany. Their cub remains, and incomplete chewed prey bones from mammoths and woolly rhinoceroses, typical of hyena activity, underline the character of these sites as cub-raising and communal dens, where their prey was accumulated along the riverbanks in a similar manner to modern African hyenas.  相似文献   

4.
Here, we present and discuss results from geo‐archaeological and palaeo‐zoological investigations at the Palaeolithic site Pymva Shor, in the Russian Arctic. As many as 3324 vertebrate fauna remains were recovered during two excavations. This includes bones of mammals, birds and fish. Radiocarbon dates were obtained from 26 specimens. The results show ages in the range 30–3 cal. ka BP. Hare and reindeer are the best represented amongst the identified mammalian species, whilst ptarmigan and various wader species dominate the avian bones. The Pleistocene assemblage includes herbivorous herd animals such as horse, bison and musk ox. These species are typical of the treeless tundra‐steppe landscape that existed during the Lateglacial. Of particular interest is a cave lion specimen that has been radiocarbon dated to approximately 15.5 cal. ka BP. According to our knowledge, this is one of the latest dated examples of this species in Eurasia. The faunal composition in the Holocene assemblage is strikingly different and includes distinct forest taxa such as beaver and pine marten. The avifauna also supports a forested environment with the presence of black grouse. A few stone artefacts were found within the strata, and have been radiocarbon dated to 16–15 cal. ka BP, suggesting that there were humans in the Pymva Shore area at that time. We identified impact notches and cut marks on some radiocarbon‐dated reindeer and bison bones, showing that humans were present twice during the Younger Dryas period. A fourth occupation phase is identified during the mid‐Holocene (6–5 cal. ka BP). We also investigated river terraces and obtained a series of luminescence dates. These have been used to reconstruct the geological history and the relationship to the find‐bearing strata.  相似文献   

5.
Dawson tephra, recently recognized in the Klondike area of Yukon Territory, records one of the largest Quaternary volcanic eruptions in Beringia. Its composition is similar to that of Old Crow tephra, indicating a source in the Aleutian arc-Alaska Peninsula region of southwestern Alaska. Its primary thickness in central Yukon is nearly twice that of Old Crow tephra, which has an estimated eruption volume of >50 km3. The distribution of Dawson tephra is still poorly known, but based on its source area and occurrence in central Yukon, it should be widespread across southern Alaska, Yukon and the Gulf of Alaska. New radiocarbon ages indicate the eruption occurred at about 24,000 14C yr BP (ca 27,000 cal yr BP). The Dawson tephra is a valuable marker bed for correlating late Pleistocene records across large areas of eastern Beringia and adjacent marine records.  相似文献   

6.
Fossil arctic ground squirrel (Spermophilus parryii) middens were recovered from ice-rich loess sediments in association with Sheep Creek-Klondike and Dominion Creek tephras (ca 80 ka) exposed in west-central Yukon. These middens provide plant and insect macrofossil evidence for a steppe-tundra ecosystem during the Early Wisconsinan (MIS 4) glacial interval. Midden plant and insect macrofossil data are compared with those previously published for Late Wisconsinan middens dating to ～25–2914C ka BP (MIS 3/2) from the region. Although multivariate statistical comparisons suggest differences between the relative abundances of plant macrofossils, the co-occurrence of steppe-tundra plants and insects (e.g., Elymus trachycaulus, Kobresia myosuroides, Artemisia frigida, Phlox hoodii, Connatichela artemisiae) provides evidence for successive reestablishment of the zonal steppe-tundra habitats during cold stages of the Late Pleistocene. Arctic ground squirrels were well adapted to the cold, arid climates, steppe-tundra vegetation and well-drained loessal soils that characterize cold stages of Late Pleistocene Beringia. These glacial conditions enabled arctic ground squirrel populations to expand their range to the interior regions of Alaska and Yukon, including the Klondike, where they are absent today. Arctic ground squirrels have endured numerous Quaternary climate oscillations by retracting populations to disjunct “interglacial refugia” during warm interglacial periods (e.g., south-facing steppe slopes, well-drained arctic and alpine tundra areas) and expanding their distribution across the mammoth-steppe biome during cold, arid glacial intervals.  相似文献   

7.
渤海湾西岸BT113孔35ka以来的沉积环境演化与海陆作用   总被引：1，自引：0，他引：1  
根据渤海湾西岸有孔虫和介形虫与年代学(AMS14C和OSL测年)研究,将该段岩心自下而上划分为6个沉积单元(U1—U6),它们依次形成于晚更新世的河流及全新世的潮滩、浅海、前三角洲、三角洲前缘和三角洲平原沉积环境,结束于约35 ka的U1单元的陆相环境,证实晚更新世低海面时渤海湾西岸未受海水影响。U1和U2单元之间,存在历时约27 ka的沉积间断(35～8.5 ka cal BP),研究区因此缺失早全新世沉积。U2单元的潮滩环境指示全新世海侵于8.5 ka cal BP到达渤海湾西岸,当时相对海平面为-16.7 m。U3单元为浅海环境,约6 ka cal BP时相对海平面上升到-6.8～-1.8 m。8.5～6 ka cal BP期间的海面上升速率是0.4～0.6 cm/a,可能与MWP1C事件有关。U4—U5单元,转为三角洲环境,沉积速率增大,反映自3.7 ka cal BP河流输入影响加强,约1.3 ka cal BP时成陆,形成U6单元。晚更新世35 ka以来的陆海环境演化,总体上是对气候变化控制下的海面变化的响应。  相似文献   

8.
Late Quaternary glacier fluctuations in the Macha Khola valley (Gorkha Himal, Nepal) were reconstructed using relative and absolute dating techniques. Our results indicate that younger moraine complexes were left by Late Holocene (<1.7 cal. ka BP), mid-Holocene (ca 3 cal. ka BP), and Lateglacial (ca 13 cal. ka BP) ice advances. Older Late Quaternary glacier advances occurred during Marine Oxygen Isotope Stages (MIS) 2 and 3–4. No relics of Middle or Early Pleistocene glaciations could be found. During MIS 3–4, glaciers advanced down to an altitude of at least 2150 m a.s.l., corresponding to an ELA depression of approximately 1300 m. At about 3500 m a.s.l., the MIS 2 Macha Khola glacier reached almost the thickness of the former MIS 3–4 glacier and retreated some time before 17.9 cal. ka BP. The Lateglacial glacier advanced again several times to altitudes between 2450 and 3400 m a.s.l. The mid-Holocene glaciers extended much farther down-valley than the Late Holocene ones. Dendrochronological data of Abies spectabilis suggested several periods of unfavourable growth conditions especially at the beginning of the 19th (1820) and 20th (1905) centuries.  相似文献   

9.
Unglaciated parts of the Yukon constitute one of the most important areas in North America for yielding Pleistocene vertebrate fossils. Nearly 30 vertebrate faunal localities are reviewed spanning a period of about 1.6 Ma (million years ago) to the close of the Pleistocene some 10 000 BP (radiocarbon years before present, taken as 1950). The vertebrate fossils represent at least 8 species of fishes, 1 amphibian, 41 species of birds and 83 species of mammals. Dominant among the large mammals are: steppe bison (Bison priscus), horse (Equus sp.), woolly mammoth (Mammuthus primigenius), and caribou (Rangifer tarandus) – signature species of the Mammoth Steppe fauna (Fig. 1), which was widespread from the British Isles, through northern Europe, and Siberia to Alaska, Yukon and adjacent Northwest Territories. The Yukon faunas extend from Herschel Island in the north to Revenue Creek in the south and from the Alaskan border in the west to Ketza River in the east. The Yukon holds evidence of the earliest-known people in North America. Artifacts made from bison, mammoth and caribou bones from Bluefish Caves, Old Crow Basin and Dawson City areas show that people had a substantial knowledge of making and using bone tools at least by 25 000 BP, and possibly as early as 40 000 BP. A suggested chronological sequence of Yukon Pleistocene vertebrates (Table 1) facilitates comparison of selected faunas and indicates the known duration of various taxa.  相似文献   

10.
Changes in the ostracod assemblages from two sediment cores collected from Lago Cardiel in southeastern Patagonia (49°S) reflect the main regional abrupt climatic changes over the last 15.6 cal. ka BP. Shifts in species abundance and switches in dominances suggest that these were mainly driven by variable salinity. During the Late Pleistocene, Limnocythere rionegroensis was abundant and dominant, indicating waters with high salinity and prevalence of evaporative processes. Between 12.6 and 10.8 cal. ka BP, Lago Cardiel expanded markedly and reached an Early Holocene highstand of +55 m above present lake level. A major change in ostracod assemblage in which Limnocythere patagonica appears as the dominant species in parallel with the disappearance of L. rionegroensis and Eucypris aff. cecryphalium mirrored this transitional period between the cold and dry Late Pleistocene and the humid and warm Early Holocene. Over the last 4 cal. ka BP, L. rionegroensis returned to the species assemblage and Riocypris whatleyi increased its abundance pointing towards increasing salinities. The variations in size, shape and ornamentation of L. rionegroensis and R. whatleyi fossil valves were examined using geometric morphometric techniques and further compared to those of modern Patagonian sites. Limnocythere rionegroensis specimens displayed high morphological variability during the evolution of Lago Cardiel. More specifically, the switch in reproductive mode – from sexual to parthenogenetic– and the increase in valve ornamentation around 12.7 cal. ka BP suggest that these changes were promoted by the hydrological alteration that occurred in the Late Pleistocene. This exercise provides a robust range of morphological variation for these proxies, which will be useful in further taxonomic and palaeoenvironmental studies adding more information about different factors influencing the observed morphological trends.  相似文献   

11.
Global climate change at the end of Pleistocene led to extinction in the huge territories of Northern Eurasia of the typical representatives of the Mammoth fauna: mammoth, woolly rhinoceros, wild horse, bison, musk-ox, and cave lion. Undoubtedly the Mammoth fauna underwent pressure from Upper Paleolithic humans, whose hunting activity could also have played a role in decreasing the number of mammoths and other representatives of megafauna. Formerly it was supposed that the megafauna of the “Mammoth complex” had become extinct by the beginning of the Holocene. Nevertheless the latest data indicate that extinction of the Mammoth fauna was significantly delayed in the north of Eastern Siberia. In the 1990s some radiocarbon dates established that mammoths existed in the Holocene on Wrangel Island—from 7700 until 3700 yBP. Radiocarbon data show that wild horses inhabited the north of Eastern Siberia 4600–2000 yBP. Muskoxen lived here about 3000 yBP. Some bison remains from Eastern Siberia belong to the Holocene. The following circumstances could promote the survival of representatives of Mammoth fauna. Cool and dry climate in this region promotes the maintenance of steppe associations—the habitats of those mammals. Late Paleolithic and Mesolithic settlements are not found in the Arctic zone of Eastern Siberia from Taimyr Peninsula to the lower Yana River; they are very rare in basins of the Indigirka and Kolyma Rivers. The small number of Stone Age hunting tribes in the northern part of Eastern Siberia was probably another factor that contributed to the survival of some Mammoth fauna representatives.  相似文献   

12.
Plant and insect macrofossil assemblages dating to the full-glacial (late Wisconsinan) are rare from eastern Beringia. Here we present an assemblage of fossil pollen, insect and plant macrofossils recovered from alluvium at the Bluefish Exposure, northern Yukon Territory. Nine AMS radiocarbon ages place these data between ca. 18,880–16,440 14C yr BP (22,313–19,597 cal. yr BP). These data indicate that xeric steppe, rich in bunchgrasses Poa and Elymus, Artemisia frigida and diverse forbs was interspersed within a mosaic of local vegetation types, including mid-rich fens, mesic graminoid meadows, steppe-tundra and herb-tundra. Macrofossils and minor pollen of tundra forbs suggest steppe-tundra plant associations within midslope elevations and discontinuous herb-tundra on high elevation uplands on exposed bedrock ridges. The composition and distribution of local vegetation was dependent on available moisture, drainage, aspect and elevation. Compositional and physiognomic similarities can be made with extrazonal steppe-dominated dry slopes and high elevation steppe-tundra ecotones in central Alaska and Yukon Territory. Our paleoecological data reflect environments inhabited by the diverse late Pleistocene Bluefish Caves fauna, including woolly mammoth, horse, steppe bison, and saiga antelope.  相似文献   

13.
The Moringa Cave within Pleistocene sediments in the En Gedi area of the Dead Sea Fault Escarpment contains a sequence of various Pleistocene lacustrine deposits associated with higher-than-today lake levels at the Dead Sea basin. In addition it contains Chalcolithic remains and 5th century BC burials attributed to the Persian period, cemented and covered by Late Holocene travertine flowstone. These deposits represent a chain of Late Pleistocene and Holocene interconnected environmental and human events, echoing broader scale regional and global climate events. A major shift between depositional environments is associated with the rapid fall of Lake Lisan level during the latest Pleistocene. This exposed the sediments, providing for cave formation processes sometime between the latest Pleistocene (ca. 15 ka) and the Middle Holocene (ca. 4500 BC), eventually leading to human use of the cave. The Chalcolithic use of the cave can be related to a relatively moist desert environment, probably related to a shift in the location of the northern boundary of the Saharo-Arabian desert belt. The travertine layer was U-Th dated 2.46 ± 0.10 to 2.10 ± 0.04 ka, in agreement with the archaeological finds from the Persian period. Together with the inner consistency of the dating results, this strongly supports the reliability of the radiometric ages. The 2.46-2.10 ka travertine deposition within the presently dry cave suggests a higher recharge of the Judean Desert aquifer, correlative to a rising Dead Sea towards the end of the 1st millennium BC. This suggests a relatively moist local and regional climate facilitating human habitation of the desert.  相似文献   

14.
《Quaternary Science Reviews》2007,26(17-18):2031-2035
Recent debate about the timing of late Pleistocene extinctions in North America has taken place on the radiocarbon timescale. Since the current internationally agreed radiocarbon calibration curve (known as IntCal04) extends back well into the Pleistocene, it is possible to make inferences on the calendar scale. To do so requires some fairly sophisticated, tailored statistical tools, to allow for (a) the presence of considerable uncertainty on individual radiocarbon ages and on the IntCal04 estimate, and (b) the inevitable incompleteness of our access to the fossil record. In this paper we demonstrate Bayesian radiocarbon calibration software, known as BCal, which implements models with both of these features, is tried and tested within the archaeology research community, but has not previously been used by those engaged in extinction research. We conclude that the extinction of horse (Equus ferus/caballus) in Alaska and Yukon is broadly contemporary with the arrival of humans in the area and took place at around 14,200 cal BP. We find that the extinction of mammoth (Mammuthus primigenius) in the same region occurred around 900 calendar years later (c. 13,300 cal BP). We also establish, with high probability, that the start of the Bölling warm phase occurred before these events and that the start of the Younger Dryas cold phase occurred after.  相似文献   

15.
These data indicate that the last representatives of Hyaenidae on the Oka plateau, i.e., cave hyenas (C. c. spelaea), are direct descendants of the hyenas that lived in the Altai–Sayan and Baikal mountain country from the Late Miocene to the end of the Pleistocene. Their continued existence over a long time undoubtedly indicates the absence of repeated glaciation during the Pleistocene. Cave hyenas are relics of the late Tertiary fauna in the area of Lake Baikal; they became extinct due to gradual cooling, which peaked at the end of the Late Pleistocene.  相似文献   

16.
The cave bear was a prominent member of the Upper Pleistocene fauna in Eurasia. While breakthroughs were recently achieved with respect to its phylogeny using ancient DNA techniques, it is still challenging to date cave bear fossils beyond the radiocarbon age range. Without an accurate and precise chronological framework, however, key questions regarding the palaeoecology cannot be addressed, such as the extent to which large climate swings during the last glacial affected the habitat and possibly even conditioned the final extinction of this mammal. Key to constraining the age of cave bear fossils older than the lower limit of radiocarbon dating is to date interlayered speleothems using 230Th/U. Here we report new results from one such site in the Eastern European Alps (Schwabenreith Cave), which yielded the highest density of bones of cave bear (Ursus spelaeus eremus). Although dating of the flowstones overlying this fossiliferous succession was partly compromised by diagenetic alteration, the 230Th/U dates indicate that the bear hibernated in this cave after about 113 ka and before about 109 ka. This time interval coincides with the equivalent of Greenland Stadial 25, suggesting possible climate control on the cave bear's habitat and behaviour. © 2019 The Authors. Journal of Quaternary Science Published by John Wiley & Sons Ltd  相似文献   

17.
The fossil vole Microtus deceitensis occurs in Early Pleistocene deposits at Fort Selkirk, Yukon Territory, and Late Pliocene beds at the type locality, Cape Deceit, Alaska. Analyses of simple vs complex morphotypes in the cheek teeth, and of differentiation of tooth enamel, show that the Cape Deceit sample of M. deceitensis is less derived, and thus appears to be older, than the Fort Selkirk sample. The fossiliferous deposits at Fort Selkirk are well constrained by fission-track and radiometric dates and are 1.5 to 1.7 myr. Sediments at Cape Deceit bear a normal magnetic polarity, are correlated with the Olduvai subchron, and probably are latest Pliocene.  相似文献   

18.
Glacial landforms and outwash terraces in the Nenana River valley, Reindeer Hills and Monahan Flat in the central Alaska Range were dated with 60 10Be exposure ages to determine the timing of Late Pleistocene glaciation. In the Nenana River valley, glaciation occurred at 104–180 ka (Lignite Creek glaciation), ca. 55 ka (Healy glaciation), and ca. 16 ka (Carlo Creek phase); glaciers retreated in the Reindeer Hills and Monahan Flat by ca. 14 ka and ca. 13 ka, respectively. The Carlo Creek moraine is similar in age to at least six other moraines in the Alaska Range, Ahklun Mountains and Brooks Range. The new data suggest that post‐depositional geological processes limit the usefulness of 10Be methods to the latter part (≤60 ka) of the late Quaternary in central Alaska. Ages on Healy and younger landforms cluster well, with the exception of Riley Creek moraines and Monahan Flat‐west sites, where boulders were likely affected by post‐depositional processes. Copyright © 2010 John Wiley & Sons, Ltd.  相似文献   

19.
The cave bear ( Ursus spelaeus ) was one of several spectacular megafaunal species that became extinct in northern Eurasia during the late Quaternary. Vast numbers of their remains have been recovered from many cave sites, almost certainly representing animals that died during winter hibernation. On the evidence of skull anatomy and low δ15N values of bone collagen, cave bears appear to have been predominantly vegetarian. The diet probably included substantial high quality herbaceous vegetation. In order to address the reasons for the extinction of the cave bear, we have constructed a chronology using only radiocarbon dates produced directly on cave bear material. The date list is largely drawn from the literature, and as far as possible the dates have been audited (screened) for reliability. We also present new dates from our own research, including results from the Urals. U. spelaeus probably disappeared from the Alps and adjacent areas – currently the only region for which there is fairly good evidence – c . 24 000 radiocarbon years BP ( c . 27 800 cal. yr BP), approximately coincident with the start of Greenland Stadial 3 ( c . 27 500 cal. yr BP). Climatic cooling and inferred decreased vegetational productivity were probably responsible for its disappearance from this region. We are investigating the possibility that cave bear survived significantly later elsewhere, for example in southern or eastern Europe.  相似文献   

20.
An AMS radiocarbon-dated pollen record from a peat deposit on Mitkof Island, southeastern Alaska provides a vegetation history spanning ∼12,900 cal yr BP to the present. Late Wisconsin glaciers covered the entire island; deglaciation occurred > 15,400 cal yr BP. The earliest known vegetation to develop on the island (∼12,900 cal yr BP) was pine woodland (Pinus contorta) with alder (Alnus), sedges (Cyperaceae) and ferns (Polypodiaceae type). By ∼12,240 cal yr BP, Sitka spruce (Picea sitchensis) began to colonize the island while pine woodland declined. By ∼11,200 cal yr BP, mountain hemlock (Tsuga mertensiana) began to spread across the island. Sitka spruce-mountain hemlock forests dominated the lowland landscapes of the island until ∼10,180 cal yr BP, when western hemlock (Tsuga heterophylla) began to colonize, and soon became the dominant tree species. Rising percentages of pine, sedge, and sphagnum after ∼7100 cal yr BP may reflect an expansion of peat bog habitats as regional climate began to shift to cooler, wetter conditions. A decline in alders at that time suggests that coastal forests had spread into the island's uplands, replacing large areas of alder thickets. Cedars (Chamaecyparis nootkatensis, Thuja plicata) appeared on Mitkof Island during the late Holocene.  相似文献   

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