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1.
Comparing the Performance of Forest gap Models in North America   总被引：6，自引：0，他引：6  
Forest gap models have a long history in the study of forest dynamics, including predicting long-term succession patterns and assessing the potential impacts of climate change and air pollution on forest structure and composition. In most applications, existing models are adapted for the specific question at hand and little effort is devoted to evaluating alternative formulations for key processes, although this has the potential to significantly influence model behavior. In the present study, we explore the implications of alternative formulations for selected ecological processes via the comparison of several gap models. Baseline predictions of forest biomass, composition and size structure generated by several gap models are compared to each other and to measured data at boreal and temperate sites in North America. The models ForClim and LINKAGES v2.0 were compared based on simulations of a temperate forest site in Tennessee, whereas FORSKA-2V, BOREALIS and ForClim were compared at four boreal forest sites in central and eastern Canada. Results for present-day conditions were evaluated on their success in predicting forest cover, species composition, total biomass and stand density, and allocation of biomass among species. In addition, the sensitivity of each model to climatic changes was investigated using a suite of six climate change scenarios involving temperature and precipitation. In the temperate forest simulations, both ForClim and LINKAGES v2.0 predicted mixed mesophytic forests dominated by oak species, which is expected for this region of Tennessee. The models differed in their predictions of species composition as well as with respect to the simulated rates of succession. Simulated forest dynamics under the changed climates were qualitatively similar between the two models, although aboveground biomass and species composition in ForClim was more sensitive to drought than in LINKAGES v2.0. Under a warmer climate, the modeled effects of temperature on tree growth in LINKAGES v2.0 led to the unrealistic loss of several key species. In the boreal forest simulations, ForClim predicted significant forest growth at only the most mesic site, and failed to predict a realistic species composition. In contrast, FORSKA-2V and BOREALIS were successful in simulating forest cover, general species composition, and biomass at most sites. In the climate change scenarios, ForClim was highly sensitive, whereas the other two models exhibited sensitivity only at the drier central Canadian sites. Although the studied sites differ strongly with respect to both the climatic regime and the set of dominating species, a unifying feature emerged from these simulation exercises. The major differences in model behavior were brought about by differences in the internal representations of the seasonal water balance, and they point to an important limitation in some gap model formulations for assessing climate change impacts.  相似文献   

2.
The forest succession model FORSKA was applied to a west-east transect across Central Europe using points from a global climate data set. Climate change experiments were undertaken for two general circulation model scenarios and two different site classes. The simulated climate changes lead to reduced forest productivity and a changed species composition on most sites. Under current climate, the broad scale pattern of the climatically driven distribution of forest communities is quite realistically reproduced. However, the resolution of climate data imposes limitations on the simulation of forest dynamics in subcontinental climate, because climate variability and extreme events are not well represented.  相似文献   

3.
The forest model ForClim was used to evaluate the applicability of gap models in complex topography when the climatic input data is provided by a global database of 0.5° resolution. The analysis was based on 12 grid cells along an altitudinal gradient in the European Alps. Forest dynamics were studied both under current climate as well as under four prescribed 2 × CO2 scenarios of climatic change obtained from General Circulation Models, which allowed to assess the sensitivity of mountainous forests to climatic change.Under current climate, ForClim produces plausible patterns of species composition in space and time, although the results for single grid cells sometimes are not representative of reality due to the limited precision of the climatic input data.Under the scenarios of climatic change, three responses of the vegetation are observed, i.e., afforestation, gradual changes of the species composition, and dieback of today's forest. In some cases widely differing species compositions are obtained depending on the climate scenario used, suggesting that mountainous forests are quite sensitive to climatic change. Some of the new forests have analogs on the modern landscape, but in other cases non-analog communities are formed, pointing at the importance of the individualistic response of species to climate.The applicability of gap models on a regular grid in a complex topography is discussed. It is concluded that for their application on a continental scale, it would be desirable to replace the species in the models by plant functional types. It is suggested that simulation studies like the present one must not be interpreted as predictions of the future fate of forests, but as means to assess their sensitivity to climatic change.  相似文献   

4.
The continuous provisioning of forest ecosystem goods and services (EGS) is of considerable interest to society. To provide insights on how much EGS provision will change with a changing climate and which factors will influence this change the most, we simulated forest stands on six climatically different sites in Central Europe under several scenarios of species diversity, management, and climate change. We evaluated the influence of these factors on the provision of a range of tree-based EGS, represented by harvested basal area, total biomass, stand diversity, and productivity. The most influential factor was species diversity, with diverse forest stands showing a lower sensitivity to climate change than monocultures. Management mainly influenced biomass, with the most intensively managed stands retaining more of their original biomass than others. All three climate-change scenarios yielded very similar results. We showed that (1) only few factor combinations perform worse under climate-change conditions than others, (2) diversity aspects are important for adaptive management measures, but for some indicators, management may be more important than diversity, and (3) at locations subject to increasing drought, the future provision of EGS may decrease regardless of the factor combination. This quantitative evaluation of the influence of different factors on changes in the provision of forest EGS with climate change represents an important step towards the design of more focused adaptation strategies and highlights key factors that should be considered in simulation studies under climate change.  相似文献   

5.
We used an individual-based forest simulator (a gap model) to assess the potential effects of anthropogenic climatic change on conifer forests of the Pacific Northwestern United States. Steady-state simulations suggested that forest zones could be shifted on the order of 500–1000 m in elevation, which could lead to the local extirpation of some high-altitude species. For low-elevation sites, species which currently are more abundant hundreds of kilometers to the south would be favored under greenhouse scenarios. Simulations of transient responses suggested that forest stands could show complex responses depending on initial species composition, stand age and canopy development, and the magnitude and duration of climatic warming. Assumptions about species response to temperature, which are crucial to the model's behaviors, were evaluated using data on species temperature limits inferred from regional distributions. The high level of within-species variability in these data, and other confounding factors influencing species distributions, argue against over-interpreting simulations. We suggest how we might resolve critical uncertainties with further research.  相似文献   

6.
This study aims to demonstrate the potential of a process-based regional ecosystem model, LPJ-GUESS, driven by climate scenarios generated by a regional climate model system (RCM) to generate predictions useful for assessing effects of climatic and CO2 change on the key ecosystem services of carbon uptake and storage. Scenarios compatible with the A2 and B2 greenhouse gas emission scenarios of the Special Report on Emission Scenarios (SRES) and with boundary conditions from two general circulation models (GCMs) – HadAM3H and ECHAM4/OPYC3 – were used in simulations to explore changes in tree species distributions, vegetation structure, productivity and ecosystem carbon stocks for the late 21st Century, thus accommodating a proportion of the GCM-based and emissions-based uncertainty in future climate development. The simulations represented in this study were of the potential natural vegetation ignoring direct anthropogenic effects. Results suggest that shifts in climatic zones may lead to changes in species distribution and community composition among seven major tree species of natural Swedish forests. All four climate scenarios were associated with an extension of the boreal forest treeline with respect to altitude and latitude. In the boreal and boreo-nemoral zones, the dominance of Norway spruce and to a lesser extent Scots pine was reduced in favour of deciduous broadleaved tree species. The model also predicted substantial increases in vegetation net primary productivity (NPP), especially in central Sweden. Expansion of forest cover and increased local biomass enhanced the net carbon sink over central and northern Sweden, despite increased carbon release through decomposition processes in the soil. In southern Sweden, reduced growing season soil moisture levels counterbalanced the positive effects of a longer growing season and increased carbon supply on NPP, with the result that many areas were converted from a sink to a source of carbon by the late 21st century. The economy-oriented A2 emission scenario would lead to higher NPP and stronger carbon sinks according to the simulations than the environment-oriented B2 scenario.  相似文献   

7.
We added certain aspects of species-specific phenology, and of local frost regimes to a standard invididual-based patch model of forest stand dynamics, which we used to explore the possible consequences of four climate-change scenarios in eight distinct forest regions in British Columbia, Canada. According to model projections, lowland temperate coastal forests will be severely stressed because forest tree species will no longer have their winter-chilling requirements met. High-elevation coastal forests may either remain stable or decrease in productivity, while interior subalpine forests may eventually resemble those now found in the coastal mountains. Southern interior forests are likely to persist relatively unchanged, while boreal and sub-boreal forests of the northern interior may become dominated by Douglas-fir and western larch, rather than by spruce and pine as at present. The rate of change in forest composition may be very high in some cases. Changes under the four climate-change scenarios generally vary in magnitude but not in direction. This exercise illustrates that different forest types might respond to a changing climate for different reasons, and at different rates.  相似文献   

8.
Scaling Issues in Forest Succession Modelling   总被引：5，自引：0，他引：5  
This paper reviews scaling issues in forest succession modelling, focusing on forest gap models. Two modes of scaling are distinguished: (1) implicit scaling, i.e. taking scale-dependent features into account while developing model equations, and (2) explicit scaling, i.e. using procedures that typically involve numerical simulation to scale up the response of a local model in space and/or time. Special attention is paid to spatial upscaling methods, and downscaling is covered with respect to deriving scenarios of climatic change to drive gap models in impact assessments. When examining the equations used to represent ecological processes in forest gap models, it becomes evident that implicit scaling is relevant, but has not always been fully taken into consideration. A categorization from the literature is used to distinguish four methods for explicit upscaling of ecological models in space: (1) Lumping, (2) Direct extrapolation, (3) Extrapolation by expected value, and (4) Explicit integration. Examples from gap model studies are used to elaborate the potential and limitations of these methods, showing that upscaling to areas as large as 3000 km2 is possible, given that there are no significant disturbances such as fires or insect outbreaks at the landscape scale. Regarding temporal upscaling, we find that it is important to consider migrational lags, i.e. limited availability of propagules, if one wants to assess the transient behaviour of forests in a changing climate, specifically with respect to carbon storage and the associated feedbacks to the atmospheric CO2 content. Regarding downscaling, the ecological effects of different climate scenarios for the year 2100 were compared at a range of sites in central Europe. The derivation of the scenarios is based on (1) imposing GCM grid-cell average changes of temperature and precipitation on the local weather records; (2) a qualitative downscaling technique applied by the IPCC for central and southern Europe; and (3) statistical downscaling relating large-scale circulation patterns to local weather records. Widely different forest compositions may be obtained depending on the local climate scenario, suggesting that the downscaling issue is quite important for assessments of the ecological impacts of climatic change on forests.  相似文献   

9.
Process-based models used to investigate forest ecosystem response to climate change were not necessarily developed to include the effect of carbon dioxide (CO2) and temperature increases on physiological processes. Simulation of the impacts of climate change with such models may lead to questionable predictions. It is generally believed that significant shifts in the performance of black spruce (Picea mariana [Mill] B.S.P.) will occur under climate change. This species, which accounts for 64% of Ontario's coniferous growing stock and 80% of the annual allowable cut, represents important economic activity throughout the boreal forest region. Forest management planning requires relatively accurate productivity estimates. Thus, it is imperative to ensure that process-based models realistically predict the effect of climate change. In this study, CENTURY and FOREST-BGC models were calibrated for a productive, upland black spruce stand in northwestern Ontario. Even though both models predicted similar relative outcomes after 100 years of climate change, they disagreed on the impacts of temperature in combination with an increase in CO2. Also, absolute amounts of carbon sequestered varied with climate change scenarios. Comparison of both models indicated that the representation of critical processes in these two forest ecosystem models is incomplete. For instance, the interactive effects of CO2 and temperature increases on physiological processes at stand and soil levels are not well documented nor are they easily identifiable in the models. Their incorporation into models is therefore problematic. Practitioners must consequently be wary of assumptions about the inclusion of critical processes in models.  相似文献   

10.
Forest gap models have been used widely in the study of forest dynamics, including predicting long-term succession patterns and assessing the potential impacts of climate change on forest structure and composition. However, little effort is devoted to predict forest dynamics in the high elevation areas, although they have the sensitive response to global climate change. In the present study, based on a modified height-diameter function, we developed a new version (FAREAST-GFSM) of the forest patch model, FAREAST for simulating the changes of subalpine forests. The observed data from the Gongga Mt. Alpine Station were also used to test model precision. With the improved performance of FAREAST-GFSM, we explored the impact of three warming scenarios on subalpine forest on the eastern Tibetan plateau within a 100-year period. The study result indicates that the effects of climate change were evident on subalpine forests in the high elevation areas. The response of different species to the warming climate might eventually transform the subalpine Abies fabric forest into Betula utilis forest similar to that which is now widely distributed in the eastern Tibetan Plateau mountainous areas with the relatively lower elevation. Subalpine forests could move to higher and colder areas, which are currently tundra.  相似文献   

11.
A comparison of forest gap models: Model structure and behaviour   总被引：4，自引：0，他引：4  
Forest gap models share a common structure for simulating tree population dynamics, and many models contain the same or quite similar ecological factors. However, a wide variety of formulations are being used to implement this general structure. The comparison of models incorporating different formulations is important for model validation, for assessing the reliability of model projections obtained under scenarios of climatic change, and for the development of models with a wide range of applicability. This paper reviews qualitative and quantitative comparisons of the structure and behaviour of forest gap models.As examples of qualitative model comparisons, the different formulations used for the heightdiameter relationship, for the maximum growth equation, and for the effects of temperature and drought on tree growth are reviewed. The variety of formulations currently in use has the potential to influence simulation results considerably, but we conclude that little is known on the sensitivity of the models in this respect.The quantitative model comparisons performed so far allow us to draw the following conclusions: (1) Gap models are quite sensitive to the formulation of climate-dependent processes under current climate, and this sensitivity is even more pronounced under a changed climate. (2) Adaptations of forest gap models to specific regions have required detailed sub-models of species life history, thus complicating model comparison. (3) Some of the complex models developed for region-specific applications can be simplified without hampering the realism with which they simulate species composition. (4) Attempts to apply the models without modification beyond the area for which they were developed have produced controversial results.It is concluded that the sensitivity of forest gap models to the exact process formulations should be examined carefully, and that more systematic comparisons of model behaviour at a range of test sites would be desirable. Such studies could improve our understanding of forest dynamics considerably, and they would help to focus future research activities with gap models.  相似文献   

12.
Three gap models, KOPIDE, NEWCOP, and ForClim, were compared with respect to their structure and behavior at four sites along an elevational gradient on Changbai Mt., northeastern China, under current climate and six climate change scenarios. This study intends to compare the three gap models under identical conditions, using a standardized simulation protocol. The three models were originally developed with different backgrounds and for different purposes. While they are relatively similar in the level of structural detail they include, they still differ in many respects regarding the assumptions that are made for representing specific ecological processes.The simulations showed that none of the three gap models provides satisfactory results in all situations; each gap model has strong and weak points in its behavior. While all models are fairly successful in simulating the composition of dominant species along the gradient under current climatic conditions, their projections under a set of hypothetical scenarios of climatic change diverge rather strongly. The analysis of these simulation results shows that several problem areas need to be addressed before any of the models can be used for a reliable impact assessment.Recommendations for improvements of the models are made, including the formulation of temperature and drought effects on tree establishment and tree growth, the size of the species pool, the appropriate choice of patch size and disturbance regimes, and allometric relationships. When aiming to use gap models under new environmental conditions, we propose to carefully reconsider their formulations based on our knowledge of the relevant processes in the region under concern, instead of using the models in an `as-is' mode.  相似文献   

13.
The FORSKA2 patch model was used to simulate responses of forest biomass and species composition to four GCM projections of climate change at 11 locations along a transect oriented northeast-southwest across the boreal zone of central Canada. In agreement with earlier results, FORSKA2 produced estimates of present-day biomass accumulation and functional types very consistent with local inventory data. Simulated responses to the four GCM scenarios of climate change produced different results. The GFDL scenario consistently reduced total biomass accumulation compared to present-day conditions, whereas the other three GCMs produced overall increases. In the north, where ecosystem productivity is thought to be limited by low temperature, changes in steady-state biomass accumulation and species composition were relatively minor. In the south, where productivity is probably limited by summer water deficits, the GCM scenarios resulted in larger absolute changes, with generally large increases under GISS, and OSU and generally smaller increases under UKMO. Pronounced changes in species composition were not evident in most simulations, with the exception that warmer winter temperatures evidently allowed invasion by species currently excluded through intolerance to winter minima.  相似文献   

14.
We use a frame-based simulation model to estimate future rate of advance of the arctic treeline in response to scenarios of transient changes in temperature, precipitation, and fire regime. The model is simple enough to capture both the short-term direct response of vegetation to climate and the longer-term interactions among vegetation, fire, and insects that are important features of dynamic vegetation models. We estimate a 150–250 yr time lag in forestation of Alaskan tundra following climatic warming and suggest that, with rapid warming under dry conditions, there would be significant development of boreal grassland-steppe, a novel ecosystem type that was common during the late Pleistocene and today occurs south of the boreal forest in continental regions. Together, the time lag and grassland development would delay the positive feedback of vegetation change to climatic warming, providing a window of opportunity to control fossil fuel emissions, the primary cause of this warming.  相似文献   

15.
Climate change scenarios with a high spatial and temporal resolution are required in the evaluation of the effects of climate change on agricultural potential and agricultural risk. Such scenarios should reproduce changes in mean weather characteristics as well as incorporate the changes in climate variability indicated by the global climate model (GCM) used. Recent work on the sensitivity of crop models and climatic extremes has clearly demonstrated that changes in variability can have more profound effects on crop yield and on the probability of extreme weather events than simple changes in the mean values. The construction of climate change scenarios based on spatial regression downscaling and on the use of a local stochastic weather generator is described. Regression downscaling translated the coarse resolution GCM grid-box predictions of climate change to site-specific values. These values were then used to perturb the parameters of the stochastic weather generator in order to simulate site-specific daily weather data. This approach permits the incorporation of changes in the mean and variability of climate in a consistent and computationally inexpensive way. The stochastic weather generator used in this study, LARS-WG, has been validated across Europe and has been shown to perform well in the simulation of different weather statistics, including those climatic extremes relevant to agriculture. The importance of downscaling and the incorporation of climate variability are demonstrated at two European sites where climate change scenarios were constructed using the UK Met. Office high resolution GCM equilibrium and transient experiments.  相似文献   

16.
Recent temperature observations suggest a general warming trend that may be causing the range of tree species to shift to higher latitudes and altitudes. Since biotic interactions such as herbivory can change tree species composition, it is important to understand their contribution to vegetation changes triggered by climate change. To investigate the response of forests to climate change and herbivory by wild ungulates, we used the forest gap model ForClim v2.9.6 and simulated forest development in three climatically different valleys in the Swiss Alps. We used altitudinal transects on contrasting slopes covering a wide range of forest types from the cold (upper) to the dry (lower) treeline. This allowed us to investigate (1) altitudinal range shifts in response to climate change, (2) the consequences for tree species composition, and (3) the combined effect of climate change and ungulate herbivory. We found that ungulate herbivory changed species composition and that both basal area and stem numbers decreased with increasing herbivory intensity. Tree species responded differently to the change in climate, and their ranges did not change concurrently, thus causing a succession to new stand types. While climate change partially compensated for the reductions in basal area caused by ungulate herbivory, the combined effect of these two agents on the mix of the dominant species and forest type was non-compensatory, as browsing selectively excluded species from establishing or reaching dominance and altered competition patterns, particularly for light. We conclude that there is an urgent need for adaptive forest management strategies that address the joint effects of climate change and ungulate herbivory.  相似文献   

17.
An 11-year remotely sensed surface albedo dataset coupled with historical meteorological and stand-level forest management data for a variety of stands in Norway’s most productive logging region is used to develop regression models describing temporal changes in forest albedo following clear-cut harvest disturbance events. Datasets are grouped by dominant tree species, and two alternate multiple regression models are developed and tested following a potential-modifier approach. This result in models with statistically significant parameters (p?<?0.05) that explain a large proportion of the observed variation, requiring a single canopy modifier predictor coupled with either monthly or annual mean air temperature as a predictor of a stand’s potential albedo. Models based on annual mean temperature predict annual albedo with errors (RMSE) in the range of 0.025–0.027, while models based on monthly mean temperature predict monthly albedo with errors ranging between of 0.057–0.065 depending on the dominant tree species. While both models have the potential to be transferable to other boreal regions with similar forest management regimes, further validation efforts are required. As active management of boreal forests is increasingly seen as a means to mitigate climate change, the presented models can be used with routine forest inventory and meteorological data to predict albedo evolution in managed forests throughout the region, which, together with carbon cycle modeling, can lead to more holistic climate impact assessments of alternative forest harvest scenarios and forest product systems.  相似文献   

18.
The extensive forests of Eastern Eurasia cover an area of ca. 6 million km2. The FAREAST model, a forest gap model that simulates the stand composition and dynamics of Eastern Eurasian forests under the current climate, was used to simulate the responses of the Eastern Eurasia Forests to the climate change. Two different scenarios of possible future climatic change were obtained from the IPCC (2001) report (CMIP2 and IS92a-GS) and were used as input to the FAREAST model to determine the compositional and structural sensitivity to climate changes for several locations and along montane elevation gradients. The simulation results suggest that, under the influence of the conditions in the two climate-change scenarios, the underlying forest dynamics should be quite different. Further, Eastern Eurasian forests maintain currents forest structure and biomass only within a small range of climate change. Broad-leaved deciduous trees of such genera as Fraxinus, Quercus and Tilia increase their ranges over Eastern Eurasia under the climate-change scenarios. Conifers, such as Larix and Picea, decrease sharply under climate change and the area of their distributions are reduced. The overall biomass of Pinus is not decreased over the region. While the Pinus distribution range shifts, the area associated with the range of the taxa is not changed.  相似文献   

19.
An integrated program of ecosystem modeling and field studies in the mountains of the Pacific Northwest (U.S.A.) has quantified many of the ecological processes affected by climatic variability. Paleoecological and contemporary ecological data in forest ecosystems provided model parameterization and validation at broad spatial and temporal scales for tree growth, tree regeneration and treeline movement. For subalpine tree species, winter precipitation has a strong negative correlation with growth; this relationship is stronger at higher elevations and west-side sites (which have more precipitation). Temperature affects tree growth at some locations with respect to length of growing season (spring) and severity of drought at drier sites (summer). Furthermore, variable but predictable climate-growth relationships across elevation gradients suggest that tree species respond differently to climate at different locations, making a uniform response of these species to future climatic change unlikely. Multi-decadal variability in climate also affects ecosystem processes. Mountain hemlock growth at high-elevation sites is negatively correlated with winter snow depth and positively correlated with the winter Pacific Decadal Oscillation (PDO) index. At low elevations, the reverse is true. Glacier mass balance and fire severity are also linked to PDO. Rapid establishment of trees in subalpine ecosystems during this century is increasing forest cover and reducing meadow cover at many subalpine locations in the western U.S.A. and precipitation (snow depth) is a critical variable regulating conifer expansion. Lastly, modeling potential future ecosystem conditions suggests that increased climatic variability will result in increasing forest fire size and frequency, and reduced net primary productivity in drier, east-side forest ecosystems. As additional empirical data and modeling output become available, we will improve our ability to predict the effects of climatic change across a broad range of climates and mountain ecosystems in the northwestern U.S.A.  相似文献   

20.
Predicting future changes in tropical rainforest tree communities requires a good understanding of past changes as well as a knowledge of the physiology, ecology and population biology of extant species. Climate change during the next hundred years will be more similar to climate fluctuations that have occurred in the last few thousand years and of a much smaller magnitude than the extent of climate change experienced during last glaciation or at the Pleistocene–Holocene transition. Unfortunately, the extent to which tropical rainforest tree communities have changed during the last few thousand years has been little investigated. As a consequence we lack the detailed evidence for population and range shifts of individual tropical species resulting from climate change analogous to the evidence available for temperate zone forests. Some evidence suggests that the rate of tropical forest change in the last several thousand years may have been high. If so, then CO2 increases and the likely alterations in temperature, forest turnover rate, rainfall, or severe droughts may drive substantial future forest change. How can we predict or model the effects of climate change on a highly diverse tree community? Explanations for the regulation of tropical tree populations often invoke tree physiology or processes that are subject to physiological regulation such as herbivory, pathology or seed production. In order to incorporate such considerations into climate change models, the physiology of a very diverse tree community must be understood. My work has focused on simplifying this diversity by categorizing the shade-tolerant species into functional physiological groups. Most species and most individual trees are shade-tolerant species, gap-requiring species being relatively uncommon. Additionally, in a regenerating gap most of the individuals are shade-tolerant species that established before gap formation. Despite the fact that the shade-tolerant species are of major ecological importance, their comparative physiology has received little attention. I have found that shade-tolerant species differ substantially in their responses to light flecks, treefall light gaps and drought. Furthermore, among phylogenetically unrelated species, these differences in physiology can be predicted from leaf lifetime. These results provide a general framework for understanding the mechanics of tropical rainforests from a physiological perspective that can be used to model their responses to climate change.  相似文献   

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