共查询到20条相似文献，搜索用时 22 毫秒
1.
室内试验研究小头虫的生物扰动作用对沉积物油污降解的影响。实验在混加定量20~#柴油的小生境中进行6个月。期间对氧化还原电位不连续展(RPD)深度,Eh,烃类氧化菌菌量以及油浓度等参数做了连续测定。结果表明,小头虫的生物扰动作用通过改善沉积物次表层氧含量水平而促进了烃类氧化菌的生长,并因此使得沉积物中油污的生物降解提高了15％。 相似文献
2.
The flatfish yellowfin sole (Limanda aspera), northern rock sole (Lepidopsetta polyxystra), and Alaska plaice (Pleuronectes quadrituberculatus) in the southeastern Bering Sea prey mainly on infauna. Spatial correspondence between their stomach contents and infauna assemblages across habitat types was examined to identify indices of prey availability for flatfish habitat characterization and quality assessment. Benthic samples and flatfish stomachs were collected in 2009 near the Alaska Peninsula in the southeastern Bering Sea. Polychaetes and bivalves were the most dominant infauna groups, each comprising 35–60% by weight in each infauna sample. These two were also the only prey groups that frequently averaged > 50% of stomach content by weight. Bivalves dominated the infauna biomass on the relatively sandy inner shelf (0–50 m depth). The muddier middle shelf (50–100 m) had the highest infauna biomass, which was dominated by polychaetes. Diet compositions of the flatfish varied spatially in correspondence with the infauna assemblage. Polychaetes were prevalent in all flatfish diets on the middle shelf, even yellowfin sole whose typical primary prey are amphipods and bivalves. Polychaete-rich habitats are potentially prime for flatfish as polychaetes are readily utilized where available and generally have high nutritional value. Flatfish did not select for specific polychaete taxa, so an index of habitat quality could be based on the biomass of aggregate polychaetes or on dominant polychaete families of the region. Under normal environmental conditions, the three flatfish have slightly-offset spatial distributions, enabling each to utilize different infauna assemblages across the shelf. However, during cold phases in the Bering Sea ecosystem, as when this study was conducted, a cold pool of < 2 °C bottom water from the spring ice melt extends over the middle shelf in summer. This physiological barrier displaces all three flatfish to the inner shelf, intensifying competition for prey resources. 相似文献
3.
Abstract. The role of a cave-dwelling mysid population as a matter carrier was evaluated in a cave of the Medes Is. (NW Mediterranean) during 1988–89. Hemimysis spehmcola (Ledoyer , 1963) is a gregarious mysid whose swarms migrate daily from the inner end of the cave - where they remain during the day–to the exterior where they feed during the night. Fecal pellet composition, pellet egestion and pellet decomposition were measured in order to evaluate the transfer of matter by mysids. Based on the strong daily behavioural rhythms of these mysids. special importance was attached to the sampling frequency (every 2 h). In order to assess seasonal variability, four daily cycles were evaluated within a year. In situ incubations were carried out to determine changes on the egestion rates, morphology, weight and composition (AA. C: N ratios) of pellets in the course of a day. The number of pellets deposited inside the cave was estimated using containers distributed along the cave bottom by SCUBA divers. Fecal pellets showed an amorphous composition, whereby diatom frustules, dinoflagellate loricae and coccolithoporids were very scarce. About 25% of the body weight were estimated to be egested daily as fecal pellets, suggesting a detritivorous feeding habit. The population oscillated seasonally between 1 and 12 millions. Individuals egested between 1.6 and 3.5 pellets a day into the cave, each averaging from 9.4 to 11.9 μg DW, 0.5 to 1.0 μg C and 0.046 to 0.27 μg N. Therefore, the population carried daily about 20 407 g DW POM. 2–21 g C and 0.5 2.7 g N from outside to inside the cave. Pellets decomposed very quickly; between 20 to 50% of both C and N were released from pellets in less than 2 h after egestion. Oxidation of pellets theoretically consumes an amount of oxygen which agrees with the high BOD values previously reported for this cave. Marine caves are generally viewed as strictly oligotrophic systems; dense mysid populations, however, could strongly modify the trophic relationships in marine caves. 相似文献
4.
饵料、温度和盐度对两种海洋桡足类摄食与排粪的影响 总被引：1，自引：0，他引：1
中华哲水蚤是中国近海典型的浮游桡足类,猛水蚤是青岛胶州湾和汇泉湾常见底栖桡足类,研究2种海洋桡足类的摄食和排粪规律可为大量培养桡足类提供最佳条件。论文研究了在6种不同饵料(球等鞭金藻、亚心形扁藻、小球藻、绿色巴夫藻、三角褐指藻、小新月菱形藻)的条件下,温度和盐度对中华哲水蚤和猛水蚤摄食和排粪的影响。温度和盐度对2种桡足类摄食和排粪的影响具有种间差异性。中华哲水蚤在15℃的摄食率、滤水率和排粪率高于在20和25℃的摄食率、滤水率和排粪率。在15、20和25℃3个温度范围内,猛水蚤的摄食率、滤水率和排粪率随温度升高而升高。在20～36盐度范围内,中华哲水蚤的摄食率、滤水率和排粪率随盐度的升高先升高后降低,在盐度为28时达到最大值,猛水蚤的摄食率、滤水率和排粪率随盐度升高而逐渐降低,猛水蚤摄食和排粪的最适盐度为20。温度和盐度实验均表明,在6种饵料中,亚心形扁藻和球等鞭金藻有利于中华哲水蚤的摄食,球等鞭金藻有利于猛水蚤的摄食,而小球藻对2种桡足类的摄食均不利。 相似文献
5.
《Estuarine, Coastal and Shelf Science》2006,66(1-2):123-134
Large areas of the bottom sediments of the Baltic Sea are temporarily or permanently anoxic. These sediments are also an important sink for a variety of contaminants. Reoxygenation of bottom waters allows recolonisation by benthic infauna, which may have important implications for the fate of buried contaminants. This study used tracers to experimentally examine the role of bioturbation by benthic infauna in transporting sediment-associated contaminants in the Baltic Sea. Three different tracer methods were used in two experiments, using three key Baltic macrofaunal species: the amphipod crustacean Monoporeia affinis; the Baltic clam Macoma baltica; and the priapulid worm Halicryptus spinulosus. In the first experiment, a reoxygenation–recolonisation scenario was recreated in the laboratory, using hypoxic sediment cores collected in the field, to determine if there was remobilisation of buried 137Cs from the Chernobyl nuclear accident in 1986. The potential for the infauna to bury newly settled surface contamination was also investigated, using a fluorescent particle tracer. In the second experiment, artificially-created radiolabelled tracer layers (14C and 51Cr) were used to quantify both upward and downward movements of organic matter and sediment-associated contaminants by bioturbation.In both experiments there were clear visual differences between the sediment effects of the three species. Halicryptus spinulosus buried deepest into the sediment, creating a network of burrows, Monoporeia affinis burrowed actively in the upper few centimeters of the sediment, and Macoma baltica was quite stationary, but appeared to filter- and deposit feed at the sediment surface. Mixing depths in the hypoxic sediment varied from 4.0 ± 3.5 cm for M. baltica to 7.8 ± 2.1 cm for H. spinulosus. Biodiffusion rates (Db) were similar for all treatments but biotransport rates (r) were significantly different between treatments, mainly due to a high r value for H. spinulosus. In the experiment with radiolabelled tracer layers, 51Cr was transported more than 14C, and tracer originally at the surface transported more than tracer buried 4 cm below the surface. There was also transport of all tracers in treatments without added macrofauna. The most likely explanation is bioturbation by the meiofauna that were undoubtedly present in both experiments.Bioturbation by macrofauna both buries surface contaminants and remobilises those that are buried, but the effects are small and on a similar scale to transport caused by meiofauna. In addition, 137Cs profiles at the hypoxic site indicated that resuspension and redeposition of sediment by physical processes had occurred, and also showed that contaminants from the last 40 years were still present in the top 5–10 cm of the sediment, well within active mixing depths. At this site, as at many others in the Baltic, physical processes are likely to be far more important than biological processes in the redistribution of contaminants on a decadal timescale. 相似文献
6.
M.L. Botton 《Estuarine, Coastal and Shelf Science》1984,18(2):209-220
The intertidal flats of the Cape May, New Jersey shore of Delaware Bay are populated by large numbers of laughing gulls and migrating shorebirds during the spring and early summer. Exclusion of birds from a shallow slough and a sand bar had only minor effects on the infaunal benthic invertebrate assemblage at either site. The Cape May beaches provide a rich source of food in the form of horseshoe crab (Limulus polyphemus) eggs; foraging on this item may be more profitable than probing the sediment for infauna. Gemma gemma, a small, thick-shelled bivalve, composed over 98% of the benthic infauna at both sites in 1980, and this species may be resistant to predation by certain shorebirds, as suggested by Schneider (1978). 相似文献
7.
《Oceanologica Acta》2002,25(3-4):139-147
The annelid fauna inhabiting soft bottoms was analysed during February and April, 1997 in the Urías estuary which is mostly surrounded by mangrove swamps, but is also subjected to the effect of several anthropogenic activities. Classification and ordination techniques were used to perform the characterisation of the system based on the annelid assemblages according to environmental variables (mainly organic matter content and percentage of mud). Three zones were found: (1) ‘very polluted zone’, in front of a slaughterhouse and a fish processing factory with a total lack of macrofauna due to the strong enrichment of the sediment in organic matter; (2) ‘polluted zones’, characterised by the sporadic presence of oligochaetes in front of a domestic sewage effluent, by the dominance of Capitella capitata close to a thermoelectric plant, and by the dominance of Streblospio benedicti and oligochaetes in the surroundings of an upstream shrimp farm; (3) ‘unpolluted zones’, in the mouth of the estuary and an area surrounded by mangrove trees, with low organic matter content in sediment, high diversity and no dominant species. 相似文献
8.
Constructed wetlands, especially mangroves, have been studied for their usefulness in sewage treatment but the effects of mangrove vegetation and a sewage load on mangrove macrofauna have been given little attention. Ocypodid crabs are important components of mangrove forests and constitute good bioindicators of the functioning of the ecosystem as a whole. In constructed mangrove mesocosms, three vegetation treatments (bare substratum, and Avicennia marina and Rhizophora mucronata seedlings) were subjected to 0, 20, 60 and 100% sewage loads from a nearby hotel. The physiological condition of introduced Uca annulipes and Uca inversa was evaluated in terms of their RNA/DNA ratio after one, five and twelve months, and used as an indicator of ecological function in the system. Crab condition in 0% sewage load was similar to that of wild crabs throughout, suggesting no significant effects of the mesocosms on their RNA/DNA ratio. Overall, both species coped well with the administered sewage loads, suggesting good ecological function in the system. Both species manifested similar patterns in RNA/DNA ratio, being more affected by seasonal fluctuations than by sewage load and vegetation presence and type. Higher RNA/DNA ratios were recorded in the long compared to the short rainy season. Sewage enhanced crab condition in the bare substratum and R. mucronata treatments, especially after one year, probably as a result of enhanced food availability. Uca inversa may be more sensitive to sewage pollution than U. annulipes. In A. marina, no difference in crab condition was observed between sewage loads, and this mangrove yielded the best reduction in sewage impacts. Our results support the usefulness of constructed mangrove areas in sewage treatment, especially if planted with A. marina and inhabited by physiologically healthy ocypodid crabs to enhance the system's performance. 相似文献
9.
《Oceanologica Acta》1998,21(2):319-324
An assessment was made of the distribution of the faecal sterol coprostanol (COP) in sediments in the Bilbao estuary. Concentrations of COP ranged from 135.15 to 10.77 μg g−1 dry weight, from the inner to the open estuary, with a mean value of 51.43 μg g−1; the highest concentrations were found near the mouths of several tribularies flowing into the estuary. Comparisons with other published values indicate that sediments from the Bilbao estuary are affected by sewage pollution in a similar degree to that reported for other estuaries in the world. Organic enrichment and anaerobic sediment conditions prevail in three-fourths of its watercourse bed. Since the spatial distribution of several organic sedimentary parameters could be clearly correlated with COP levels, we can conclude the existence of major pollution by the addition of untreated sewage, coming from mainly domestic sources. The implementation of a sewerage treatment plan by the local water authority needs the monitoring of reliable sewage tracers, such as faccal sterols, to assess the effectiveness of the plan in reducing the effects of pollution. 相似文献
10.
Submerged vegetation complexity modifies benthic infauna communities: the hidden role of the belowground system 
下载免费PDF全文
下载免费PDF全文 Vanessa González‐Ortiz Luis Gonzalo Egea Rocio Jiménez‐Ramos Francisco Moreno‐Marín José Lucas Pérez‐Lloréns Tjeerd Bouma Fernando Brun 《Marine Ecology》2016,37(3):543-552
Marine plants provide a variety of functions with high economic and ecological values in ecosystems. The above‐ (AG) and below‐ground (BG) systems increase the structural complexity of plants, which also enhance faunal abundance and diversity. The ecological role of the AG compartment in structuring inter‐tidal macrobenthic communities has been widely studied; however, this is not the case for the BG compartment. This study addressed the effects of variation in vegetation complexity (in both AG and BG systems) on associated macrobenthic infauna with respect to abundance, species richness, composition, weight and body type. To achieve this aim, a field experiment using artificial vegetation mimics was carried out using replicated treatments with different AG‐BG complexity ratios. We found a significant increase in the density and the number of taxa of macrobenthic infaunal species in plots with vegetation mimics compared with unvegetated areas, regardless of either AG or BG complexity. This effect was found even when AG parts were not present (i.e. when only BG parts were used). Furthermore, a positive relationship between structural complexity and diversity was recorded. Variation in one or both plant compartments was strongly related to diversity changes in the associated macrobenthic infauna. In conclusion, our experimental set‐up provides the first evidence that the BG compartment is at least as important as the AG compartment in controlling diversity in inter‐tidal vegetated areas because it was able to strongly affect community structure even when the AG system was totally absent. 相似文献
11.
Silvana Ciarelli Wies A.P.M.A. Vonck Nico M. van Straalen 《Marine environmental research》1997,43(4):329-343
Corophium volutator (Pallas; Amphipoda: Corophiidae) is an ecologically important representative of the infauna of marine sediments and is widely distributed along the north-east American and the west European coasts. Methods for conducting short-term toxicity tests with lindane-spiked sediment using this species were developed. Corophium volutator were used in 10-d lindane-spiked sediment tests conducted on seven occasions throughout the year, to investigate the repeatability of test methods and to quantify differences in sensitivity for lindane due to seasonal field sampling. The 10-d LC50 values found for lindane ranged from 0.78 to 1.49 μg/g dry weight (dw), a relatively small range. Higher values were found in the months of April 1994 and April 1995 (1.46 and 1.49 μg/g, respectively). Positive controls conducted with cadmium (aqueous phase) also revealed lower sensitivity of amphipods in those months. The results however, show that a factor of two difference in LC50 values among tests conducted in different seasons was within the range of expected variability. A study on the size-structure of the wild amphipod population indicated that animals of all size-classes were available throughout the whole year except for the months of April and May. Experiments conducted with organisms of different length-classes revealed that differences in sensitivity to lindane were not size-related. The present study also supports the use of field-collected organisms of ecological relevance, such as C. volutator, in ecotoxicological studies. 相似文献
12.
Robert P. Griffiths Bruce A. Caldwell William A. Broich Richard Y. Morita 《Estuarine, Coastal and Shelf Science》1982,15(2):183-198
Subarctic marine sediments were exposed to fresh and ‘weathered’ crude oil from Cook Inlet, Alaska. The crude oil was thoroughly mixed with the sediment at various concentrations and some of the oil-sediment mixtures were placed on top of unamended sediments. Both sediments with added oil and those overlain by oiled sediments were investigated. These sediments were placed either in trays or aquaria. The trays were incubated in situ near the site from which the sediments were originally collected. The aquaria were maintained at simulated in situ conditions by maintaining a continuous flow of fresh seawater through them. The sediments were exposed to crude oil for periods up to 1.5 years. Sediments exposed to 50‰ fresh crude oil showed significant decreases in nitrogen fixation and denitrification rates and redox potentials. Also observed were increases in CO2 production rates and methane concentrations. These same changes were observed in sediments exposed to 1‰. When ‘weathered’ crude oil was added to the sediments, the same changes were observed except there was no reduction in nitrogen fixation activity. In most cases, the observed effects were less marked when the sediments were amended with ‘weathered’ crude oil than with the same concentration of fresh crude oil. Untreated sediments that were overlain with treated sediments showed the same changes as those sediments that were thoroughly mixed with crude oil. The presence of fresh crude oil at 50‰ essentially eliminated burrowing activity of the benthic infauna. Under these conditions, there was an accumulation of detritus particles on the oiled sediments that was not present in the untreated controls. 相似文献
13.
-Morphology, reproduction, ecology and isoenzyme electrophoresis of a Capitella complex in Qingdao were examined from 1983 to 1984. We determined that the Capitella complex in Qingdao was composed of at least three sibling species: Capitella cf capitata, Capitella sp. I and Capitella sp. IT. Although these three species are very similar in morphology, they differ in: three kinds of isoenzyme electrophoresis, staining reactions, developmental pattern, larval morphology and dispersal, reproductive strategies, breeding season and population ecology. In addition, they show a series of minor quantitative variations in adult morphology, especially in number and morphology of setae. Such differences can be detected only at the population level. Some ecological data such as density, growth rate, mortality and sex distribution are also provided in this paper. 相似文献
14.
A biological survey of the subtidal fauna near an intertidal combined sewer overflow (CSO) discharging into Puget Sound, Washington, USA, was conducted during April and May, 1977. The purpose of this study was to determine the effect of the CSO on the surrounding marine invertebrate communities.The shallowest subtidal sites studied (9 m below MLLW) appeared to be more affected by the CSO than deeper sites (13 and 24 m below MLLW). The subtidal area affected, as reflected in the levels of organic material, heavy metals and pesticides in the sediments, as well as the composition of the benthic macrofauna, appeared to be highly localised. The numerically dominant species at sites closest to the CSO was the polychaete Capitella capitata while the abundance of the bivalve Axinopsida serricata increased with distance from the CSO.Burrowing deposit feeders were the numerically dominant polychaete feeding type at sites nearest the CSO. Surface deposit feeders were more abundant at sites farther from the CSO, perhaps due to reduced sedimentation rates.Site-classification analyses indicated that sites adjacent to the CSO were more similar to one another than to sites which were farther away and apparently less affected.The positions of the sites along the first three principal component ordination axes were significantly correlated with depth and volatile organics (traceable to the CSO) in the sediments. 相似文献
15.
This study provides the first response data for ten Southern California macrophytes exposed to untreated, primary, secondary and secondary chlorinated sewage effluent during long-term culture studies in the laboratory as well as short-term metabolic studies in the field and in the laboratory. Bossiella orbigniana, Lithothrix aspergillum and Corallina officinalis var, chilensis had relatively broad homeostatic capabilities and enhanced net production rates when exposed to primary treated sewage. In the long-term cultures, Amphiroa zonata, B. orbigniana and C. officinalis var. chilensis all demonstrated enhanced growth in the presence of primary sewage. Chlorination of effluent had only a short-term negative effect; i.e. species' responses to secondary and secondary chlorinated sewage were virtually identical beyond the second week in culture. Three populations of C. officinalis var. chilensis with differing pollution histories showed a tolerance to sewage corresponding to the extent of previous exposure, indicating that this species may be able to acclimate physiologically to sewage stress. The more sewage-tolerant macrophytes displayed lower photosynthetic quotient values during exposure to effluent than those possessing little tolerance to sewage. 相似文献
16.
Morris H. Roberts 《Marine environmental research》1980,3(1):63-80
The acute toxicity of chlorinated and bromochlorinated treated sewage waste was determined with spot, Leiostomus xanthurus. The halogenated sewage waste was generated in a pilot plant located at the James River Sewage Treatment Plant, Newport News, Va. The 96 h and 144 h LC50's were 0·23 mg/l (measured residual) for chlorine and 0·25 mg/l (measured residual) for bromine chloride. These concentrations are considerably higher than those which would be produced by optimal halogenation rates for disinfection or maximum superhalogenation rates permitted by pilot plant equipment and normal effluent dilution rates. 相似文献
17.
The soft-bottom macrofauna (> 1 mm) of Tromsøysund and Sandnessund near Tromsø, northern Norway (69°40′N, 19°0′E) was sampled quantitatively in 1983, 1990 and 1992 in order to assess enrichment effects from discharges of municipal sewage and fish factory effluents. The studies comprised 30 stations between 7 and 40 m depth at varying distances from effluent outfalls. In total 395 species/taxa were recorded. The polychaetes were the most important group both with regard to the numbers of species (146) and specimens (80% of total), but bivalves and amphipods were also common. Most stations had species numbers between 40 and 100 and densities between 2000 and 8000 ind. per m2. The species assemblages could be related to organic enrichment, depth and sampling year. Evidence suggests that the organic discharges did not seriously affect the waters surrounding Tromsø, but the generally high abundances, the prevalence of polychaetes and a numerical importance of ‘opportunistic’ species (Polydora, Chaetozone, Heteromastus) might suggest that the whole area was stimulated by the organic inputs. Sharply delimited strongly enriched zones characterized by dense populations of Capitella capitata were found on shallow sites close to large outfalls in Tromsøysund. These local and restricted effects of the discharges may be related to strong tidal currents and wave action that disperse effluent components. The faunal composition remained largely stable from 1983 to 1992 at 9 revisited stations, but the number of small bivalve and crustacean taxa increased and the abundance of the dominant polychaete Polydora socialis strongly decreased. The effluent discharges were supposed to have increased from 1983 to 1992, but the faunal changes did not provide evidence of an increased enrichment of the area during the period. 相似文献
18.
E.R. McGreer 《Marine environmental research》1982,7(2):131-149
An investigation of the factors responsible for the distribution of Macoma balthica (L.) on a mudflat receiving sewage effluent was carried out in the Fraser River estuary, British Columbia. The factors examined included reduced salinity, low levels of dissolved oxygen, effluent toxicity, chlorination, substrate grain size and the effects of metal contaminated sediments. Results showed that salinity, dissolved oxygen and sediment grain size did not satisfactorily explain the distribution of M. balthica. Unchlorinated and chlorinated sewage effluent were shown to be non-toxic in laboratory tests, and these results were confirmed by 7-day in situ bioassays. The factor which appeared to determine the distribution of M. balthica was the degree of contamination of the substrate which affected the settling and survival of larval and juvenile clams. Heavy metals occurred in high concentrations in the contaminated sediments and were considered to be the most probable controlling factor. 相似文献
19.
Daniele Nizzoli Giuseppe Castaldelli Marco Bartoli David Thomas Welsh Patricia Arriaga Gomez Anna Elisa Fano Pierluigi Viaroli 《Marine Ecology》2002,23(S1):297-306
Abstract. Dark respiration rates, dissolved inorganic nitrogen (DIN) fluxes and nitrification rates were measured at two sites in the microtidal Sacca di Goro lagoon in September 2000. DIN fluxes correlated with the biomass of the dominant macrofauna species (the amphipod Corophium spp. at station Giralda and the polychaete Neanthes spp. at station Faro). Respiration (> 6 mmol O2 m−2 h−1 ) and ammonium fluxes (> 80μmol N m−2 h−1 ) were higher at station Giralda despite the lower organic matter content (4.5 %) and lower macrofauna biomass (4 g AFDW m−2 ). At both sites ammonium fluxes were significantly correlated with the biomass of the benthic infauna, but Corophium stimulated ammonium NH4 + fluxes 3-fold compared to Neanthes. The amphipod also enhanced nitrification rates (> 300 μmol N m−2 h−1 ) due to the high density of its burrows, the higher NH4 + regeneration rates and the enhanced oxygen supply to the bacteria. 相似文献
20.
F. Montserrat W. Suykerbuyk R. Al-Busaidi T.J. Bouma D. van der Wal P.M.J. Herman 《Journal of Sea Research》2011,65(1):170-181
Benthic ecosystem engineering organisms attenuate hydrodynamic or biogeochemical stress to ameliorate living conditions. Bioturbating infauna, like the lugworm Arenicola marina, determine intertidal process dynamics by maintaining the sediment oxygenated and sandy. Maintaining the permeability of the surrounding sediment enables them to pump water through the interstitial spaces, greatly increasing the oxygen availability. In a field experiment, both lugworm presence and siltation regime were manipulated to investigate to what extent lugworms are able to cope with sedimentation of increasing mud percentage and how this would affect its ecosystem engineering. Fluorescent tracers were added to experimentally deposited mud to visualise bioturbation effects on fine sediment fractions. Lugworm densities were not affected by an increasing mud percentage in experimentally deposited sediment. Negative effects are expected to occur under deposition with significantly higher mud percentages. Surface chlorophyll a content was a function of experimental mud percentage, with no effect of lugworm bioturbation. Surface roughness and sediment permeability clearly increased by lugworm presence, whereas sediment erosion threshold was not significantly affected by lugworms. The general idea that A. marina removes fine sediment fractions from the bed could not be confirmed. Rather, the main ecosystem engineering effect of A. marina is hydraulic destabilisation of the sediment matrix. 相似文献