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1.
The stable carbon isotope compositions and the stomatal parameters (stomatal density and stomatal index) of four Cheirolepidiaceae species, Brachyphyllum ningxiaensis, Brachyphyllum obtusum, Pseudofrenelopsis dalatzensis and Pseudofrenelopsis gansuensis, were analyzed to recover the late Early Cretaceous atmospheric CO2 levels. The fossil plants were collected from 5 consecutive sedimentary members of the uppermost Zhonggou Formation. Based on the stomatal data, the estimated palaeo-atmospheric CO2 concentrations in the Jiuquan Basin during the late Early Cretaceous were 1060–882 ppmv based on the carboniferous standardization and were 641–531 ppmv based on the recent standardization; the pCO2 values present at first a decreasing and then an increasing trend within the sedimentary time of the five members. The δ13Cp values based on the 21 Brachyphyllum specimens showed a large variation, which ranged from −20.98‰ to −25.69‰, with an average of −24.2‰. The values also identified a C3 photosynthetic pathway for the Brachyphyllum specimens. The predicted δ13Ca values varied from −2.1‰ to −6.38‰, with an average of −5.03‰. These two proxies were irregular within the different members; therefore, the correlation with the change in atmospheric CO2 concentrations was not significant. Moreover, a water-stressed environment was proposed based on the δ13C values of the present fossil plants, a proposal that was also supported by the previous palaeobotanical, palynological and stratigraphical evidence. In the present study, an inconsistent relationship between the stable carbon isotope and the stomata values was apparent, which most likely indicated that the stomata numbers of the plant were more sensitive to the variation in the concentration of the atmospheric CO2, whereas the δ13C values were sensitive to the moisture conditions. 相似文献
2.
《Gondwana Research》2014,25(1):214-225
Jurodidae are thought to be some of the world's most mysterious beetles and display an intriguing mixture of characters occurring in three different suborders. Hitherto, all known fossil and extant Jurodidae were extremely rare and restricted to Russian Siberia and the Far East. Here we describe two new species, Jurodes daohugouensis sp. nov. and Jurodes pygmaeus sp. nov. from the Middle Jurassic of Daohugou, China. A key to all species of Jurodidae and their research history are given, and the detailed morphology of fossil Jurodidae is also presented. Our results confirm the presence of three ocelli in fossil Jurodidae, and reveal that hind wings of J. pygmaeus sp. nov. possess well developed radial and oblong cells, and are very similar to those of extant Jurodidae (Sikhotealinia zhiltzovae). Male genitalia of fossil Jurodidae were described herein for the first time, showing a basal trilobate structure. The presence of exposed propleuron, three pairs of external trochantins, metanepisterna with ridges, primitive characters of wing venation further support the probable placement of Jurodidae in the suborder Archostemata. J. daohugouensis and J. pygmaeus are very similar to S. zhiltzovae in having the big protruding eyes, elevated median portion of frons bearing three ocelli, pubescent body, contiguous procoxae, exposed trachantion of all coxae, elytra with rows of punctures, and wing venation. Our results show that these key characters of extant Jurodidae can be traced back to the Middle Jurassic, revealing that the family has been in a period of evolutionary stasis for at least 160 million years. Furthermore, our discoveries widen the paleogeographic distribution of fossil Jurodidae from Russian Siberia to northern China. 相似文献
3.
A new species,Ginkgo huolinhensis sp.nov.,with well-preserved cuticle from the Lower Cretaceous,Huolinhe Formation of Huolinhe Coal Field in Inner Mongolia,China,was studied morphologically and anatomi... 相似文献
4.
An inclusion in Cretaceous amber from Myanmar is described as a new fossil species in the extant liverwort genus Frullania. The name Frullania pinnata is proposed for the taxon that is characterized by entire underleaves paired with a conspicuous pinnate branching pattern, two distinctive and stable morphological features. Entire underleaves are known in several extant Frullania species but had not previously been documented in Frullania in amber. The combination of morphological characters in this new fossil species is unknown in any crown group lineage of Frullania, and may very well represent a stem lineage element of the genus. This discovery is important because it expands our understanding of the diversity of Frullaniaceae in the Burmese amber forest as well as important ramifications for the phylogenetic reconstruction of extant Frullania lineages. 相似文献
5.
浙江白垩系两种掌鳞杉科化石微细结构及其古环境意义 总被引：2，自引：0，他引：2
利用扫描电镜和生物光学显微镜对采自浙江新昌镜岭馆头组的两种已绝灭的松柏类掌鳞杉科植物Pseudofrenelopsis papillosa Chow and Tsao和Pseudofrenelopsis parceramosa Watson细微结构进行了观察研究。其中P.papillosa无论从宏观形态还是表皮构造都与模式标本完全一致;而定为Pseudofrenelopsis parceramosa的标本节间角质层较为光滑,气孔排列规则,副卫细胞上未见乳突,此外该标本气孔器大小、自由叶大小、节间长度等方面都与Pseudofrenelopsis中的其他种不太相同。并利用化石P.parceramosa的气孔比率通过最近现生亲缘种法恢复了早白垩世晚期大气CO2浓度为861~1047ml/L,得出用本种恢复早白垩世晚期CO2浓度时应用石炭纪标准较为准确,说明本种可以作为环境指标。 相似文献
6.
Anne-Marie Aucour Pierre Faure Bernard Gomez Yann Hautevelle Raymond Michels Frédéric Thévenard 《Organic Geochemistry》2009,40(7):784-794
The chemical composition of Cretaceous leaf remains showing exceptionally well preserved cuticles was investigated using pyrolysis gas chromatography–mass spectrometry (Py-GC–MS) and thermally assisted hydrolysis methylation (THM)-GC–MS. Samples of Coniferales (Frenelopsis) and Ginkgoales (Nehvizdya penalveri) leaf remains were collected from freshwater and coastal marine depositional environments. Material for pyrolysis included (i) untreated leaves and cuticles obtained after extraction from mineral rock matrix and bleaching, (ii) kerogen fraction from both materials, (iii) non-hydrolysable fraction from kerogen. The THM-GC–MS data from untreated leaves and bleached cuticles show that the fossil cuticle geopolymer essentially released aliphatic components upon thermal treatment, with a dominance of fatty acids (FAs) and n-alkanes/n-alkenes. The FAs are essentially resistant to bleaching and remain after solvent extraction. They occur mainly as short chain compounds ranging from C6 to C16 and with maximum abundance at C8–C9. The n-alkanes/n-alkenes from kerogen and the non-hydrolysable residue occur mainly as short chain compounds in the range C10–C16, with the highest abundance at C9–C12. The THM-GC–MS pyrograms of the fossil cuticles differ from those of cutan from fresh living plants. They support the preservation model via polymerization of monomers derived from cutin or from unsaturated cell FAs. 相似文献
7.
A new earwig genus with a new species, Cylindopygia falcata gen. et sp. nov., is described and illustrated based on two well-preserved, nearly complete female specimens from the Lower Cretaceous, Yixian Formation in Huangbanjigou, Liaoning Province, China. C. falcata is assigned to Pygidicranidae mainly due to the following characters: head obtuse-triangular with posterior margin straight, abdomen robust, subcylindrical and densely setose, and femora compressed and carinate. The new finding represents the earliest fossil record of Pygidicranidae and the first record of Pygidicranidae in China. 相似文献
8.
One new osmylid genus and species, Burmaleon magnificus, and one new nymphid genus and species Rafaelnymphes cratoensis are described, respectively based on inclusions in the Cretaceous Burmese amber and on a compression fossil from the Crato Formation in Brazil. The nymphid Araripenymphes seldeni, from the Crato Formation, is redescribed on the basis of a new specimen, showing possible sexual dimorphism in wing coloration, a feature extremely rare among the Neuroptera. In a recently published phylogenetic analysis of the family, the attribution to the fossil taxa (versus rejection) of the larval characters proper to the modern nymphids, has a crucial impact on the resolution of the phylogeny. The compression fossils currently attributed to the Nymphidae should be revised because their wing venation alone is not really sufficient for an accurate family attribution to Nymphidae rather than to another family of Neuroptera (viz. Osmylidae). 相似文献
9.
Axelrodichthys megadromos sp. nov. is a coelacanth described based on a single specimen collected in the lower Campanian site of Ventabren motorway, Southern France. The new species is referred to the mawsoniids because of the ornamentation of the skull roof and of the proportionally wide supraorbital series. The specimen belongs to the Mawsonia-Axelrodichthys complex based on features present on the lower jaw and on the basisphenoid. The new species is referred to the genus Axelrodichthys because of proportions of its parietonasal shield and because of the arrangement of the posterior parietals relative to the supraorbitals. Autapomorphic characters, in particular on the parasphenoid, justify the inclusion of the specimen in a new species. The occurrence of a mawsoniid in the Ibero-Amorican Island that formed part of the European Archipelago in the terminal Cretaceous is an evidence of a dispersal event from the southern land masses. The occurrence of A. megadromos in the Campanian-Maastrichtian represents the last occurrence of mawsoniids worldwide, after a gap in the fossil record of about 30 million years. This belated occurrence of Axelrodichthys extends the time range of this genus to approximately 40 myr and suggests that this genus, together with its sister genus Mawsonia, were organisms with a slow morphologic evolution. 相似文献
10.
Conifers are common in the Lower Cretaceous Zhonggou Formation of Jiuquan Basin, Gansu Province, NW China. The excellent preservation of plant fossils allows detailed investigations of the leaf epidermis by light microscope (LM) and scanning electron microscope (SEM). Based on the leaf morphology and epidermal features, two Brachyphyllum species, B. ningshiaense and B. obtusum are described. They closely resemble one another in foliar morphology and epidermal features, but have different leaf apices, cuticle features, stomatal shape and the number of subsidiary cells. The specimens consist of leafy axes with imbricate, spirally arranged, scale-like leaves. Epidermal analysis shows rectangular, pentagonal or rhomboidal unspecialized epidermal cells and abaxially longitudinally arranged stomatal rows except for the marginal areas, as well as well-developed papillae, sunken stomata and Florin rings of subsidiary cells with papillae on the inner surfaces. The imbricate and spirally arranged leaves with rather thick cuticle in both species indicate an Araucariaceae (Araucaria) or Cheirolepidiaceae affinity. However, epidermal features and the presence of Classopollis pollen in the sediments indicate a cheirolepidiaceous affinity. The morphological and anatomical characters of both taxa are interpreted as adaptations to a warm seasonally and temporarily dry palaeoclimate. 相似文献
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14.
Catfish bones from Tortonian (Miocene) freshwater beds of central Argentina are here identified as pertaining to a new species of the tropical pimelodid genus Phractocephalus. The new species differs from the other recent and fossil species of the genus in skull, pectoral girdle and spine characters. The material was found in different localities near the city of Paraná, Entre Ríos Province. The bearing horizon is the so-called “Conglomerado osífero” which constitutes the lowermost beds of the fluvial Ituzaingó Formation. The aquatic vertebrate fauna occurring in the bearing bed shows a similar generic composition to several northern South American Miocene units where Phractocephalus remains were found. This report extends the range of Phractocephalus more than 2000 km to the South. The record is in agreement with higher global temperatures and putative ample hydrographic connections of the river basins in the Paraná area with the Amazon basin until at least the early late Miocene. 相似文献
15.
David R. Sussman 《Journal of South American Earth Sciences》2011,31(4):475-478
In 2007, Reguero et al. described a 2.5 Ma erethizontid dentary from the Uquía Formation in Argentina (MACN5376) and referred it to the genus Erethizon, a genus found at present only in North America. They based their generic identification on the presence in the fossil of a markedly inflected angular process. I argue in this commentary that this single trait (the angular process) in a single incomplete fossil is insufficient evidence for the Erethizon attribution, for the following reasons: 1. The trait (the inflected angular process) is variable in modern South and North American porcupines and observer bias and/or allometry may be responsible for observed differences; 2. Among fossil South American porcupines, the inflected angular process is not unique to the Uquían fossil; 3. The fossil possesses other traits (including one trait newly described in this paper) associated with modern South American (Coendou) porcupines and not modern Erethizon; and 4. The traits by which we recognize modern Erethizon, primarily related to the genus’s ability to survive severe winters, are most readily explained by evolution driven by the South American porcupine immigrants’ exposure in North America to that severe weather. The Uquían fossil is not Erethizon and should be attributed for the present to the genus Coendou. 相似文献
16.
Multiple foliar specimens of the Late Pennsylvanian fossil pteridosperm [gymnosperm] Alethopteris zeilleri (Ragot) Wagner were collected from one restricted stratigraphical horizon in the Canadian Sydney Coalfield. Variability of functional-group distribution using FTIR technique was studied in compressions, adaxial versus abaxial cuticles, and in unseparated cuticles as a function of maceration time from 48 to 168 h. The results obtained document spectral variability that could be expected within specimens of one species. For example, CH2/CH3 and Al/ox ratios can differ by as much as 20% of the values. Moreover, the experiments performed confirm that by using a previously established maceration protocol, long maceration periods do not bias FTIR spectra in terms of oxygenation overprinting. The inference that this cuticle is robust, under the given diagenetic level, probably reflects a reassuring degree of chemical fidelity of the Pennsylvanian plant to support Carboniferous chemotaxonomic observations. 相似文献
17.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight. 相似文献
18.
《地学前缘(英文版)》2020,11(5):1875-1888
The fossil trunks and rhizomes of Osmundaceae provide important information about its evolutionary history. Due to limited records of the Mesozoic trunks and rhizomes in the Eurasia of the Northern Hemisphere, our understanding on the fossil diversity of the Osmundaceae is hindered. Two new species of the Osmundaceae trunks, Osmundacaulis asiatica sp. nov. and Osmundacaulis sinica sp. nov. representing the first discovery of the Mesozoic tree fern genus Osmundacaulis in Eurasia, are described from Wudalianchi and Qiqihar, Heilongjiang Province, Northeast China, which enriches the plant diversity of the Osmundaceae in the Eurasia, and provides vital evidence for studying the distribution, radiation and evolution of the genus during the Cretaceous. The fossil records suggest that Osmundacaulis species may have evolved from a common ancestor, which first appeared in the Australian portion of Pangaea, and then spread to ancient northern North America and ancient East Asia. Since then, they developed into different species through their own evolutionary lines. The Chinese species have a special local feature that the outer cortex is thicker than the inner cortex, in contrast with reported Osmundacaulis species having thinner outer cortex and thicker inner cortex. Long-term geographic isolation may have led to the radiation of diverse Osmundacaulis species and the appearance of region-specific features, such as the thick outer cortex and the thin inner cortex of the Chinese species. Among all reported Osmundacaulis species, the two new species found in China, O. nerii from the Jurassic of Australia and the O. lemonii from the Jurassic of the USA, have special groups of mucilage-sacs inside sclerenchyma ring of petiole base. Mucilage sacs probably originated independently among taxonomic groups, representing convergent adaptations to similar habitats, rather than indicating genetic inheritance from a common ancestor. 相似文献
19.
One new genus with three new species of the family Prionoglarididae, Palaeosiamoglaris leinhardi Azar, Huang et Nel gen. et sp. nov.; P. burmica Azar, Huang et Nel sp. nov. and P. inexpectata Azar, Huang et Nel sp. nov. are characterised, illustrated and described from Cretaceous Burmese amber. Palaeosiamoglaris gen. nov. share several characters with the recent genus Siamoglaris Lienhard, 2004, endemic to the Oriental (Sino-Indian) Region. A new tribe Siamoglaridini Azar, Huang et Nel trib. nov. is established to accommodate the group Siamoglaris + Palaeosiamoglaris; the other Prionoglaridinae being attributed to Prionoglaridini. This discovery implicates a palaeobiogeographic repartition for this particular tribe (Siamoglarini) that is still the same currently. Most diagnostic features show a nearly stasis state, supporting the notion that these characters appeared at least by the early Cenomanian. A checklist of all prionoglaridids is given. 相似文献
20.
A new genus, Cretasyne gen. nov., in the family Lasiosynidae, with two species C. lata sp. nov. and C. longa sp. nov., is described from the Lower Cretaceous Yixian Formation of Inner Mongolia, China. These fossils are the first record of the family from the Yixian Formation and provide significant information on lasiosynid morphology and diversity. The fossil record of Lasiosynidae is also reviewed. 相似文献