Speed bumps and barricades in the carbon cycle: substrate structural effects on carbon cycling     

C. Arnosti   《Marine Chemistry》2004,92(1-4):263

The observation that a fraction of organic matter produced in marine systems evades the concerted efforts of microbial communities and is buried in sediments suggests that there are ‘speed bumps’ in carbon degradation pathways that impede microbially driven remineralization processes. The initial step in degradation of macromolecules, extracellular enzymatic hydrolysis, is often stated to be ‘the’ rate-limiting step in carbon remineralization. Experimental investigations described here, however, demonstrate that at least in certain cases, microbes produce extracellular enzymes on time scales of hours to tens of hours in response to substrate addition, and hydrolysis is extremely rapid. If enzymatic hydrolysis can be rapid, what factors slow or stop organic matter degradation? A lack of the correct inducer to initiate enzyme production, and/or a lack of the correct organism to produce the required enzyme, may result in a complete lack of hydrolysis in certain environments—a barricade, rather than a speed bump. Preliminary evidence supporting this hypothesis includes a comparison of polysaccharide hydrolysis in seawater and sediments, which demonstrates that the spectrum of enzymes active in seawater and sediments are fundamentally different. Furthermore, a survey of enzyme activities in surface waters from a range of locations suggests that pelagic microbial communities also differ widely in their abilities to express specific extracellular enzymes. Trans-membrane transport through porins is yet another potential location of structure-related selectivity.Our efforts to identify speed bumps and barricades are hampered by our inability to structurally characterize in sufficient detail the macromolecular structures present in marine systems. Furthermore, assessments of organic matter ‘quality’ from a chemical perspective do not necessarily accurately reflect the availability of organic carbon to microbial communities. For these communities, in fact, ‘quality’ may be a variable, which depends on the enzymatic and uptake capabilities of community members. To begin to assess substrate structure and quality from a microbial perspective, we will have to combine specific knowledge of macromolecular structures with detailed investigations of the enzymatic and transport capabilities of heterotrophic marine microbes.  相似文献   

Møre Margin: Crustal structure from analysis of Expanded Spread Profiles     

Ingi Olafsson  Eirik Sundvor  Olav Eldholm  Kjersti Grue 《Marine Geophysical Researches》1992,14(2):137-162

Analysis in both the x—t and —p domains of high-quality Expanded Spread Profiles across the Møre Margin show that many arrivals may be enhanced be selective ray tracing and velocity filtering combined with conventional data reduction techniques. In terms of crustal structure the margin can be divided into four main areas: 1) a thicker than normal oceanic crust in the eastern Norway Basin; 2) expanded crust with a Moho depth of 22 km beneath the huge extrusive complex constructed during early Tertiary breakup; 3) the Møre Basin where up to 13–14 km of sediments overlie a strongly extended outer part with a Moho depth at 20 km west of the Ona High; and 4) a region with a 25–27 km Moho depth between the high and the Norwegian coast. The velocity data restricts the continent-ocean boundary to a 15–30 km wide zone beneath the seaward dipping reflector wedges. The crust west of the landward edge of the inner flow is classified as transitional. This region as well as the adjacent oceanic crust is soled by a 7.2–7.4 km s^–1 lower crustal body which may extend beneath the entire region that experienced early Tertiary crustal extension. At the landward end of the transect a 8.5 km s^–1 layer near the base of the crust is recognized. A possible relationship with large positive gravity anomalies and early Tertiary alkaline intrusions is noted.  相似文献   

三疣梭子蟹肝脏的结构研究   总被引：25，自引：0，他引：25  

李太武 《海洋与湖沼》1996,27(5):471-475

于１９８９年９月－１９９０年８月，用电镜研究浙江舟山出产的三疣梭子蟹肝脏的结构。结果表明，三疣梭子蟹的肝脏主要由４种细胞构成；Ｂ细胞，大，长柱状，胞质中有许多液泡，粗面内质网发达；Ｒ细胞和Ｆ细胞，两者相似，均含丰富的粗面内质网，核糖体、脂滴、线粒体等结构，但Ｒ细胞脂滴数量多，体积大，Ｆ细胞的核糖体数量超过Ｒ细胞２；Ｅ细胞，多边形，核大而圆，粗面内质网少，线粒体小而狭长。  相似文献   

新生相形成的模拟实验研究     

雍国平  陈邦林 《海洋科学》1996,20(2):37-39

通过模拟实验制备了硅酸盐类不溶性固体，从Ｘ射线衍射特征和红外光谱特征两方面看，制得的硅酸盐类与河口新生相成分，结构相似，表明铁水解成的β－ＦｅＯＯＨ胶体吸附Ｓｉ，Ａｌ，Ｍｇ，Ｋ等元素，经结构重组可形成硅酸盐矿物，这可能是河口体系新生相的形成原因。  相似文献   

云南永胜宝坪浅源热液改造型铜矿地质及成因     

谭玉荣  秦培林  王学武  高建国 《云南地质》2008,27(3):357-361

宝坪铜矿赋存于三叠系下统腊美组中。矿体呈似层状,少量脉状,严格受北北西-北西向构造控制。矿床成因类型属浅源热液改造型铜矿床。  相似文献   

Late Paleozoic volcanism in the central part of the Southern Permian Basin (NE Germany, W Poland): facies distribution and volcano-topographic hiati     

Marion Geißler  Christoph Breitkreuz  Hubert Kiersnowski 《International Journal of Earth Sciences》2008,97(5):973-989

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渤海湾盆地埕岛油田缝洞型潜山油气藏构造及储层特征   总被引：1，自引：0，他引：1  

付兆辉  陈发景  刘忠胜  张卫平  程明  张在振 《海相油气地质》2008,13(1):37-44

埕岛油田古潜山油气藏为典型的缝洞型碳酸盐岩油气藏。埕岛地区经历了多期次的构造运动,并经受了多期次的断裂与剥蚀作用,为油气成藏创造了良好条件。埕岛油田古潜山可以划分为三个构造单元,即埕北20断层下降盘的西排山,上升盘的中排山和东排山,三个潜山带成帚状敛于凸起最南端,为几个二级构造单元交汇处,构造应力较发育。埕岛油田有三种储集空间类型:晶间孔隙、溶蚀孔洞和裂缝。  相似文献   

Mt. Melbourne volcano,antarctica: Evidence of seismicity related to volcanic activity     

Salvatore Gambino  Eugenio Privitera 《Pure and Applied Geophysics》1996,146(2):305-318

Characterization of the microseismic activity (M _L <2.0) has been performed at Mt. Melbourne since 1990. We recorded a group of low frequency events with common morphological characteristics, i.e., an emerging onset, an unclear second phase and a sharply dropping coda. Spectral analysis of events recorded at more than one station indicates that the seismogram characteristics and spectral content are largely due to source effects. A polarization filter applied to a set of three component data revealed a first phase made up ofP waves followed (after about 0.9–1.4 sec.) by a second phase probably composed ofSH-type waves. Particle motion analysis detected a seismic ray angle direction mainly between N70°E and N110°E and apparent angle of incidence between 35° and 48° for the first phase. The studied seismicity was localized in an area on the eastern slope of Mt. Melbourne Volcano which presents a surface temperature anomaly (Mazzarini andSalvini, 1994). We formulate two hypotheses for the type of earthquakes recorded: 1) long-period events involving active presence of magmatic fluids in the source processes; 2) or the result of fracturing processes (shear?) in a medium characterized by transition between brittle and plastic behaviors. In the latter hypothesis the superficial thermal anomaly may be a symptom of this behavior at depth and is confirmed by the lown values observed for the exponential fit in the codaQ analysis.  相似文献   

The dynamics and thermodynamics of large ash flows   总被引：6，自引：6，他引：0  

Marcus I. Bursik  Andrew W. Woods 《Bulletin of Volcanology》1996,58(2-3):175-193

 Ash flow deposits, containing up to 1000 km³ of material, have been produced by some of the largest volcanic eruptions known. Ash flows propagate several tens of kilometres from their source vents, produce extensive blankets of ash and are able to surmount topographic barriers hundreds of metres high. We present and test a new model of the motion of such flows as they propagate over a near horizontal surface from a collapsing fountain above a volcanic vent. The model predicts that for a given eruption rate, either a slow (10–100 m/s) and deep (1000–3000 m) subcritical flow or a fast (100–200 m/s) and shallow (500–1000 m) supercritical flow may develop. Subcritical ash flows propagate with a nearly constant volume flux, whereas supercritical flows entrain air and become progressively more voluminous. The run-out distance of such ash flows is controlled largely by the mass of air mixed into the collapsing fountain, the degree of fragmentation and the associated rate of loss of material into an underlying concentrated depositional system, and the mass eruption rate. However, in supercritical flows, the continued entrainment of air exerts a further important control on the flow evolution. Model predictions show that the run-out distance decreases with the mass of air entrained into the flow. Also, the mass of ash which may ascend from the flow into a buoyant coignimbrite cloud increases as more air is entrained into the flow. As a result, supercritical ash flows typically have shorter runout distances and more ash is elutriated into the associated coignimbrite eruption columns. We also show that one-dimensional, channellized ash flows typically propagate further than their radially spreading counterparts. As a Plinian eruption proceeds, the erupted mass flux often increases, leading to column collapse and the formation of pumiceous ash flows. Near the critical conditions for eruption column collapse, the flows are shed from high fountains which entrain large quantities of air per unit mass. Our model suggests that this will lead to relatively short ash flows with much of the erupted material being elutriated into the coignimbrite column. However, if the mass flux subseqently increases, then less air per unit mass is entrained into the collapsing fountain, and progressively larger flows, which propagate further from the vent, will develop. Our model is consistent with observations of a number of pyroclastic flow deposits, including the 1912 eruption of Katmai and the 1991 eruption of Pinatubo. The model suggests that many extensive flow sheets were emplaced from eruptions with mass fluxes of 10⁹–10¹⁰ kg/s over periods of 10³–10⁵ s, and that some indicators of flow "mobility" may need to be reinterpreted. Furthermore, in accordance with observations, the model predicts that the coignimbrite eruption columns produced from such ash flows rose between 20 and 40 km. Received: 25 August 1995 / Accepted: 3 April 1996  相似文献   

华南海西—印支期成矿地质背景          下载免费PDF全文

陈志明 《地质科学》1993,28(1):61-67

华南海西—印支成矿期内,成矿序列完整,类型齐全,矿种繁多,为其它成矿期所莫及。沉积矿床需要稳定的地质背景,层控矿床形成的较有利的地质背景是稳中有动。  相似文献