Sex‐related differences in size,function and regeneration of the major cheliped in the hermit crab Pagurus filholi          下载免费PDF全文

Kento Matsuo  Daisuke Tanikawa  Chiaki I. Yasuda  Satoshi Wada 《Marine Ecology》2015,36(4):1391-1399

Pagurus hermit crabs have a well‐developed right cheliped (major cheliped) and in some species the major cheliped of males is longer than that of females. This paper describes sex‐related differences in major cheliped length and regeneration pattern of the major cheliped in the hermit crab Pagurus filholi. We also examined the function of the major cheliped in male–male competition. Major cheliped length of males was longer than that of females in P. filholi. Males regenerated larger chelipeds than females at the first molt after experimentally induced autotomy. Body size growth in males of the regeneration group was less than that in intact males of the control group while there was no significant difference in body size growth of females between regeneration and control groups. Major cheliped length was included in the best model to explain the outcome of male–male competition and thus sexual selection appears to be a causative factor in the sex‐related difference of the major cheliped length. Sex‐related differences in the regeneration pattern may reflect differences in evolutionary pressures on males for large major chelipeds and females for large body size.  相似文献   

Observations of current and mixing around the shelf break in Pribilof Canyon in the Bering Sea     

T. Tanaka  I. Yasuda  H. Onishi  H. Ueno  M. Masujima 《Journal of Oceanography》2015,71(1):1-17

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Estimates of non-tidal exchange transport between the Sea of Okhotsk and the North Pacific     

Katsuro Katsumata  Ichiro Yasuda 《Journal of Oceanography》2010,66(4):489-504

The outflow from the Sea of Okhotsk to the North Pacific is important in characterising the surface-to-intermediate-depth water masses in the Pacific Ocean. The two basins are separated by the Kuril Islands with numerous straits, among which the Bussol and the Kruzenshterna Straits are deeper than 1000 m. The physics governing the transport between the two basins is complicated, but when the semidiurnal and diurnal tides are subtracted, the observed density and velocity structures across the Bussol Strait suggest a significant contribution from geostrophic balance. Using a two-layer model with the interface at 27.5σ _θ , part of the upper layer transport that is not driven by tides is estimated using two previously unexplored data sets: outputs from the Ocean General Circulation Model for Earth Simulator (OFES), and historical hydrographic data. The Pacific water flows into the Sea of Okhotsk through the northeastern straits. The greatest inflow is through the Kruzenshtern Strait, but the OFES results show that the contributions from other shallower straits are almost half of the Kruzenshtern inflow. Similarly, the outflow from the Sea of Okhotsk is through the southwestern straits of the Kuril Islands with the largest Bussol Strait contributing 60% of the total outflow. The OFES and hydrographic estimates agree that the exchange is strongest in February to March, with an inflow of about −6 to −12 Sv (negative indicates the flow from the North Pacific, 1 Sv = 10⁶ m³s⁻¹), and an outflow from the Sea of Okhotsk of about +8 to +9 Sv (positive indicates the flow from the Sea of Okhotsk), which is weakest in summer (−3 to +1 Sv through the northeastern straits and +0 to +3 Sv through the southwestern straits). The estimated seasonal variation is consistent with a simple analytic model driven by the difference in sea surface height between the two basins.  相似文献   

Uptake mechanism of anthropogenic CO<Subscript>2</Subscript> in the Kuroshio Extension region in an ocean general circulation model     

Hideyuki Nakano  Hiroyuki Tsujino  Mikitoshi Hirabara  Tamaki Yasuda  Tatsuo Motoi  Masao Ishii  Goro Yamanaka 《Journal of Oceanography》2011,67(6):765-783

The uptake mechanism of anthropogenic CO₂ in the Kuroshio Extension is examined by a Lagrangian approach using a biogeochemical model embedded in an ocean general circulation model. It is found that the uptake of anthropogenic CO₂ is caused mainly by the increase of pCO₂ dependency of seawater on temperature, which is caused by greater dissolved inorganic carbon concentration in the modern state than in the pre-industrial state. In contrast with the view of previous studies, the effect of the vertical entrainment, which brings waters that last contacted the atmosphere with the past lower CO₂ concentration, is comparatively small. Winter uptake of anthropogenic CO₂ increases with the rise of the atmospheric CO₂ level, while summer uptake is relatively stable, resulting in a larger seasonal cycle of the uptake. This increase is significant, especially in the Kuroshio Extension region. It is newly suggested that this increase in the Kuroshio Extension region is largely caused by the combined effects of the increased pCO₂ dependency of the sea water on the temperature and the seasonal difference in cooling.  相似文献   

Effects of high-order nonlinear interactions on unidirectional wave trains   总被引：2，自引：0，他引：2  

Nobuhito Mori  Takashi Yasuda 《Ocean Engineering》2002,29(10):1233-1245

Numerical simulations of gravity waves with high-order nonlinearities in two-dimensional domain are performed by using the pseudo spectral method. High-order nonlinearities more than third order excite apparently chaotic evolutions of the Fourier energy in deep water random waves. The high-order nonlinearities increase kurtosis, wave height distribution and H_max/H_1/3 in deep water and decrease these wave statistics in shallow water. Moreover, they can generate a single extreme high wave with an outstanding crest height in deep water. High-order nonlinearities (more than third order) can be regarded as one cause of freak waves in deep water.  相似文献   

Structure of Subsurface Intrusion of the Oyashio Water into the Kuroshio Extension and Formation Process of the North Pacific Intermediate Water     

Kuniaki Okuda  Ichiro Yasuda  Yutaka Hiroe  Yugo Shimizu 《Journal of Oceanography》2001,57(2):121-140

In order to understand the actual formation process of the North Pacific Intermediate Water (NPIW), structure of subsurface intrusions of the Oyashio water and the mixing of the Oyashio and the Kuroshio waters in and around the Kuroshio Extension (KE) were examined on the basis of a synoptic CTD observation carried out in May-June 1992. The fresh Oyashio water in the south of Hokkaido was transported into KE region through the Mixed Water Region (MWR) in the form of subsurface intrusions along two main paths. The one was along the east coast of northern Japan through the First Branch of the Oyashio (FBO) and the other along the eastern face of a warm streamer which connected KE with a warm core ring through the Second Branch of the Oyashio (SBO). The fresh Oyashio water extended southward through FBO strongly mixed with the saline NPIW transported by the Kuroshio in the south of Japan (old NPIW) in and around the warm streamer. On the other hand, the one through SBO well preserved its original properties and extended eastward beyond 150°E along KE with a form of rather narrow band. The intrusion ejected Oyashio water lens with a diameter of 50–60 km southward across KE axis and split northward into the MWR involved in the interaction of KE and a warm core ring, which were supposed to be primary processes of new NPIW formation.  相似文献   

Vertical structure of the tidal current ellipse in a rotating basin     

Hidekazu Yasuda 《Journal of Oceanography》1987,43(5):309-318

Tidal current ellipses formed by the Coriolis effect are investigated theoretically, taking account of the effect of horizontal boundaries. This study reveals that even in a narrow bay, the tidal current ellipse appears in the inside of the bottom boundary layer, although the ellipticity cannot be recognized in the outside of the boundary layer. In a wide bay, the ellipticity is observed even in the outside and it is larger in the inside. The rotation direction of these ellipses is counterclockwise in the northern hemisphere. And also, the Coriolis force has an effect to deflect the major axis of the ellipse in the inside of the boundary layer. These nature of the tidal current ellipse is well explained by the consideration of the formation mechanism of the ellipse.  相似文献   

Tide-induced residual currents with Stokes and Ekman layers over an undulatory sea floor     

Hidekazu Yasuda  J. T. F. Zimmerman 《Journal of Oceanography》1986,42(4):276-293

Residual currents induced by the tidal current flowing over a rotating ridge, which is infinitely elongated in they-direction and the width of which is so large that the nonlinear effect is weak enough, have been analysed theoretically. The currents are restrained by Stokes and Ekman layers which are given through a constant viscosity. This study shows that the direction of the cross-isobath residual current just above the sea floor varies with the value off ^*(=Coriolis parameter/tide frequency), that the along-isobath residual current forms a clockwise circulation around the rotating ridge and that it is stronger asf ^* grows larger.  相似文献   

Tide-induced vertical circulation in a bay with homogeneous water: Theory and experiment     

Hidekazu Yasuda 《Journal of Oceanography》1981,37(2):74-86

The structure of the tidal residual current due to vertical viscosity is investigated both theoretically and experimentally. It is found that the interaction between the vertical component of the oscillatory current and vorticity,wT ₁ T ₁, induces a strong residual constituent outside the boundary layer and forms a circulation which is quite similar to gravitational circulation and that the vertical profile of the oscillatory current not only affects the magnitude of the residual constituent but also decides the direction of the circulation. In the hydraulic experiment, the residual constituent is larger than the theoretical prediction and a phase difference in the oscillatory constituent between the upper and lower layer is observed. The amplitude difference is caused by the strong nonlinear effect of the residual constituent and the phase difference is caused by the interaction between the residual current and the basic oscillatory current. The principal generating force of the residual constituent outside the boundary layer,wT ₁ T ₁, is observed in a bay where the tide is nearly a standing wave.  相似文献   

Orbital evolution and accretion of protoplanets tidally interacting with a gas disk: I. Effects of interaction with planetesimals and other protoplanets     

Junko Kominami  Hidekazu Tanaka 《Icarus》2005,178(2):540-552

We have performed N-body simulations on the stage of protoplanet formation from planetesimals, taking into account so-called “type-I migration,” and damping of orbital eccentricities and inclinations, as a result of tidal interaction with a gas disk without gap formation. One of the most serious problems in formation of terrestrial planets and jovian planet cores is that the migration time scale predicted by the linear theory is shorter than the disk lifetime (10⁶-10⁷ years). In this paper, we investigate retardation of type-I migration of a protoplanet due to a torque from a planetesimal disk in which a gap is opened up by the protoplanet, and torques from other protoplanets which are formed in inner and outer regions. In the first series of runs, we carried out N-body simulations of the planetesimal disk, which ranges from 0.9 to 1.1 AU, with a protoplanet seed in order to clarify how much retardation can be induced by the planetesimal disk and how long such retardation can last. We simulated six cases with different migration speeds. We found that in all of our simulations, a clear gap is not maintained for more than 10⁵ years in the planetesimal disk. For very fast migration, a gap cannot be created in the planetesimal disk. For migration slower than some critical speed, a gap does form. However, because of the growth of the surrounding planetesimals, gravitational perturbation of the planetesimals eventually becomes so strong that the planetesimals diffuse into the vicinity of the protoplanets, resulting in destruction of the gap. After the gap is destroyed, close encounters with the planetesimals rather accelerate the protoplanet migration. In this way, the migration cannot be retarded by the torque from the planetesimal disk, regardless of the migration speed. In the second series of runs, we simulated accretion of planetesimals in wide range of semimajor axis, 0.5 to 2-5 AU, starting with equal mass planetesimals without a protoplanet seed. Since formation of comparable-mass multiple protoplanets (“oligarchic growth”) is expected, the interactions with other protoplanets have a potential to alter the migration speed. However, inner protoplanets migrate before outer ones are formed, so that the migration and the accretion process of a runaway protoplanet are not affected by the other protoplanets placed inner and outer regions of its orbit. From the results of these two series of simulations, we conclude that the existence of planetesimals and multiple protoplanets do not affect type-I migration and therefore the migration shall proceed as the linear theory has suggested.  相似文献