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1.
Bathymodiolus属贻贝是广泛分布于全球深海冷泉和热液生态系统的优势种,在深海化能生态系统的物质循环和能量流动中起着重要作用。本文以我国南海北部冷泉的主要优势种平端深海偏顶蛤(Bathymodiolus platifrons)为研究对象,采用常规生化测定方法,研究了其主要生化成分和氨基酸组成的特征,并就特定组织中主要生化成分的含量与生活在热液区的平端深海偏顶蛤和近岸的远东偏顶蛤(Modiolus kurilensis)进行了对比。结果表明,南海冷泉平端深海偏顶蛤软体部含水量84.28%,粗蛋白含量7.18%,粗脂肪含量1.23%,糖类含量2.75%,与已报道的深海贝类组成相近。虽然主要生化成分含量在3种贻贝的鳃、外套膜、闭壳肌和消化腺4个组织中总体差异不大,但是冷泉平端深海偏顶蛤和热液平端深海偏顶蛤的外套膜糖类含量(25.20%、30.66%)显著高于远东偏顶蛤(6.97%,P0.05),这表明平端深海偏顶蛤的主要储存物质为外套膜中的糖类。在氨基酸组成上,冷泉平端深海偏顶蛤鳃中氨基酸总量为44.55%(干质量),外套膜中为34.83%(干质量),其中必需氨基酸分别占比41.73%和40.52%,总体与其他贝类相似。然而,在平端深海偏顶蛤中,与渗透压调节相关的甘氨酸和与硫代谢相关的牛磺酸含量较高,这与其适应深海高盐度高硫化氢浓度的环境相关。综上所述,南海冷泉平端深海偏顶蛤在常规生化组分和氨基酸组成上与近岸常见双壳类具有一定差异,这些差异与其特殊生境的关系还需要更加深入的研究。  相似文献   

2.
The vestimentiferan tubeworm Ridgeia piscesae is an ecosystem-structuring organism in the hydrothermal vent environments of the Northeast Pacific. During this study, a single representative aggregation of the long-skinny morphotype of R. piscesae from the main endeavor segment was monitored for 3 yr before being collected in its entirety with a hydraulically actuated collection device manipulated in situ by a research vehicle. Vestimentiferan growth rates in this aggregation were determined by staining the exterior of the tubes and measuring newly deposited tube sections. The average growth rate of R. piscesae in this aggregation was very low in both years of the growth study (3.2 mm yr−1). Although the incidence of plume damage from partial predation was very high (>95%), mortality was very low (<4% yr−1). The distribution and the very tight clustering of recently recruited individuals indicated gregarious settlement behavior that is hypothesized to be partly due to biotic cues from settled larvae. Coupled measurements of vent fluid sulfide concentration and temperature were used to calculate the exposure of the vestimentiferans to sulfide from short- and long-term temperature monitoring. Plume-level temperature records indicate that most of the time individuals in this aggregation were exposed to extremely low levels of vent fluid, and therefore sulfide (<0.1 μM), while their posterior sections were consistently exposed to sulfide concentrations in the 100 μM range. A rootball-like structure formed the common base of the aggregation. In contrast to the anterior sections of the tubeworm tubes, the portions of the tubes within the “rootball” were freely permeable to sulfide. The results of this study show that R. piscesae, unlike vestimentiferans from the East Pacific Rise, can survive and grow in areas of low diffuse vent flow with very low plume-level exposure to sulfide. We propose that this morphotype of R. piscesae has the ability to acquire sulfide from sources near their posterior ends, similar to some species of cold seep vestimentiferans from the Gulf of Mexico. The ability of this single species of vestimentiferan to survive low exposure to vent flow with low mortality coupled with sulfide uptake across posterior tube sections may help explain the occurrence of a single vent vestimentiferan species in a wide variety of habitat conditions at hydrothermal vent sites in the Northeast Pacific.  相似文献   

3.
Phylogenetic relationship among three species of deep-sea vesicomyid bivalvesCalyptogena, i.e.,C. soyoae, C. solidissima, C. fausta and an undescribed species of the Iheya Ridge, Okinawa Trough was analyzed on the basis of the RFLP analysis of the fragment (about 1 kbp) of cytochrome oxidase I. Both the two populations ofC. soyoae (off Hatsushima and Okinoyama Bank, Sagami Bay) consisted of two haplotypes, which could be discriminated by only one restriction site (1.7% sequence divergence).Calyptogena of the Iheya Ridge could not be distinguished from one of the two haplotypes ofC. soyoae. Nucleotide substitution rates between species were calculated and dendrograms were constructed.  相似文献   

4.
The San Clemente cold seep lies within 100–200 km of other reducing habitats in the NE Pacific, offering an opportunity to compare diversity and species overlap among reducing habitats (i.e. whale‐, kelp‐, and wood‐falls) at similar depths within a single region. Video observations from the research submersible Alvin at the San Clemente seep (1800 m depth) indicated clumps (‘thickets’) of vestimentiferans distributed as meter‐scale patches interspersed with vesicomyid clam beds and black sediments. Sediment‐core samples were collected at distances of 0 to 80–200 m along randomly oriented transects radiating outward from vestimentiferan thickets to evaluate changes in macrofaunal community structure from thickets into the background community. Macrofaunal abundance was elevated at distances of 0–1 m compared to 80–200 m (i.e. the ‘background’ community). The tube‐building frenulate worms Siboglinum spp., along with peracarid crustaceans, dominated sediments within 1 m of vestimentiferan thickets. Species diversity was depressed within 1 m of thickets but with high rates of species accumulation, suggesting that seep sites greatly increase sediment heterogeneity and facilitate colonization by non‐background macrofaunal species. Stable isotope data indicate chemosynthetic nutrition for some dominant macrofaunal species within 1 m of tubeworm thickets. The macrofaunal community near vestimentiferan thickets in San Clemente seep contains intermediate levels of species richness and diversity compared to other deep‐sea seep areas in the northeast Pacific. There was low species overlap between the San Clemente seep macrofauna and communities in reducing habitats near wood‐, whale‐, and kelp‐falls at similar depths within the region, suggesting that seeps harbor a distinct infaunal community.  相似文献   

5.
《Oceanologica Acta》2002,25(5):227-241
ChEss is a recently-funded Census of marine life programme aimed at improving our knowledge of the biogeography of deepwater chemosynthetically driven ecosystems by promoting an international field phase of discovery and exploration. The main objectives are to assess and explain the diversity, distribution and abundance of hydrothermal vent and cold seep species. With the global mid-ocean ridge system extending ∼65 000 km, it is unlikely that its entire length would be examined in detail. The ChEss programme proposes to select a limited number of target areas chosen for the discovery of new vents and seeps. The intention is to identify the maximum scientific return that could be achieved from detailed investigation of the minimum number of sites at key locations. To narrow the field for exploration, a number of starting hypotheses and goals have been identified. A bio- and geo-referenced database for hydrothermal vent and cold seep species will be created. This database will be integrated with the Ocean Biogeographic Information System (OBIS). An international scientific committee will coordinate the programme, facilitate collaboration between participants, promote ship-time applications at national level and stimulate scientific innovation from a wider community.  相似文献   

6.
软体动物腹足纲分类学研究进展——从近海到深海   总被引：2，自引：0，他引：2  
随着科学技术的不断进步和研究的不断深入,我国关于海洋的研究已由浅海逐步向深海发展,不断取得新的发现和认知。自2014年4月起,中国科学院海洋研究所依托先进的"科学"号海洋科学综合考察船,搭载"发现"号水下机器人（remotelyoperated vehicle,ROV）,先后对冲绳海槽热液区、南海冷泉区、雅浦海山区以及马努斯热液区等深海区域进行了综合性科学考查,采集到了大量深海腹足纲（Gastropoda）标本,其中有一些为未经发现和描述的新种,极大地推动了我国关于深海软体动物物种多样性研究,实现了深海环境和软体动物资源的新认知。本文对近年来从深海采集到的腹足类标本进行了整理分类,对其中的11属13种进行描述,并对深海特殊环境中腹足纲物种多样性进行了总结,对中国软体动物腹足纲从近海至深海的分类现状进行研究论述。  相似文献   

7.
Four specimens of the thyasirid Conchocele bisecta (Conrad) and one small specimen of the vesicomyid Hubertschenckia ezoensis (Yokoyama), each with a drill hole made by a naticid gastropod, were found at a cold‐seep site in the upper Eocene Poronai Formation of Hokkaido. This is apparently the oldest record not only of drill holes, but also of predation scars, in a cold‐seep fauna. In addition, drilled vesicomyids are known from several Miocene cold‐seep sites in Japan. We suggest that the Eocene and Miocene chemoautotrophic bivalves were drilled only in the shallow‐water settings preferred by most naticids. The lack of drill holes in Oligocene chemoautotrophic bivalves in the northwestern USA suggests that this innovation, which allowed naticids to prey upon highly toxic bivalves, first appeared in the western Pacific during the Eocene.  相似文献   

8.
Atomic absorption spectroscopy (flame and graphite furnace techniques) and instrumental neutron activation analysis were used for determining the Fe, Mn, Zn, Cu, Pb, Cd, Ag, Co, Cr, As, Se, Sb, Ba, Au, and Hg contents in 25 samples of different tissues and whole organisms inhabiting the southern trough in the Guaymas Basin (Gulf of California) and in several samples of its bottom waters. It is shown that the habitat environment of this hydrothermal field with high primary production of both photosynthetic and bacterial chemosynthetic origin influences the Fe and Mn ratios in the waters of the microbiotopes and the distribution patterns of the metals in the external and internal organs of the benthic animals. In the dominant specialized taxa, the maximal bioaccumulation of metals is registered both in the organs related to bacterial chemosynthesis such as the trophosome of Vestimentifera Riftia pachyptila and the gills of the vesicomyid clam Archivestica gigas and in other organs. The other organisms such as the mollusks Nuculana grasslei, actinias Paraphelliactis pabista, Actinaria, and Spongia and the crabs Munidopsis alvisca demonstrate high bioaccumulation properties as well. The metal concentration coefficient is highly variable ranging from 10 to 104. The changes in the molar Fe/Mn ratio values imply the partitioning of these two metals in the following migration succession: microbiotope water-external organs-internal organs.  相似文献   

9.
基因组大小(或称C值)作为生物单倍体细胞中全套染色体的DNA总量,在一定程度上是恒定的,因而C值可以作为生物物种的一个特定参数。深海热液和冷泉为更好地理解C值与不同环境之间的关系提供了一个特征性的模型。本文采用流式细胞术,测定了来自热液和冷泉环境中的10种深海无脊椎动物的C值,其分布范围从0.87 pg到12.28 pg,其中,相比于软体动物和多毛类,甲壳生物基因组大小及变异均较大。对比热液和冷泉两个群落中共有种(深海偏顶蛤Bathymodiolus platifrons、柯氏潜铠虾Shinkaia crosnieri以及长角阿尔文虾Alvinocaris longirostris)的基因组大小,发现C值差异并不显著。同时,综合已有的数据,对深海化能极端环境与其他环境条件下的物种C值进行对比分析,结果显示深海化能极端环境下生物的基因组大小并没有发现明确的变化趋势。  相似文献   

10.
Cold seep communities discovered at three previously unknown sites between 600 and 1000 m in Monterey Bay, California, are dominated by chemoautotrophic bacteria (Beggiatoa sp.) and vesicomyid clams (5 sp.). Other seep-associated fauna included galatheid crabs (Munidopsis sp.), vestimentiferan worms (Lamellibrachia barhami?), solemyid clams (Solemya sp.), columbellid snails (Mitrella permodesta, Amphissa sp.), and pyropeltid limpets (Pyropelta sp.). More than 50 species of regional (i.e. non-seep) benthic fauna were also observed at seeps. Ratios of stable carbon isotopes (δ13C) in clam tissues near 36‰ indicate sulfur-oxidizing chemosynthetic production, rather than non-seep food sources, as their principal trophic pathway. The “Mt Crushmore” cold seep site is located in a vertically faulted and fractured region of the Pliocene Purisima Formation along the walls of Monterey Canyon ( 635 m), where seepage appears to derive from sulfide-rich fluids within the Purisima Formation. The “Clam Field” cold seep site, also in Monterey Canyon ( 900 m) is located near outcrops in the hydrocarbon-bearing Monterey Formation. Chemosynthetic communities were also found at an accretionary-like prism on the continental slope near 1000 m depth (Clam Flat site). Fluid flow at the “Clam Flat” site is thought to represent dewatering of accretionary sediments by tectonic compression, or hydrocarbon formation at depth, or both. Sulfide levels in pore waters were low at Mt Crushmore (ca 0.2 mM), and high at the two deeper sites (ca 7.011.0 mM). Methane was not detected at the Mt Crushmore site, but ranged from 0.06 to 2.0 mM at the other sites.  相似文献   

11.
The vesicomyid Calyptogena kilmeri is one of the most abundant bivalves inhabiting chemosynthetic environments shallower than c. 1500 m along central California. We estimated the population size structure, biomass, rates of individual growth, somatic production, and mortality for C. kilmeri, based on sampling of seep habitats and tag–recapture studies at chemosynthetic communities in Monterey Bay, California. The composite growth rate of C. kilmeri over all sites was relatively high (K = 0.25), reaching c. 80% of asymptotic length (104.7 mm) in 6.6 years. The density of C. kilmeri was estimated as 938.5 ind. m−2 and biomass density varied from 704 to 2059 g ash free dry mass (AFDM) m−2. Somatic production was also high (294–297 g·AFDM·m−2 year−1), and production/biomass ratios for C. kilmeri varied from 0.14 to 0.42 among sites, related to variation in size–frequency distributions among sites. Instantaneous mortality rates estimated from size distributions ranged from 0.17 to 0.24 year−1. Growth and somatic production by C. kilmeri are in the range reported for chemosynthetic bivalves from hydrothermal vent and seep habitats, as well as photosynthetic‐based assemblages of inshore or intertidal bivalves, and greatly exceed rates reported for heterotrophic deep‐sea benthos.  相似文献   

12.
Deep faunal assemblages were observed on cold seeps, between 1000 and 2000m depth, along the southern termination of the Barbados accretionary prism, during the Caracolante II and Diapicar cruises. Faunal composition and microdistribution of a cold seep community were analyzed at one site using deep-sea photographs and seismic (3.5 Kz) profiles. The community is dominated by large mussels up to 20cm long (possibly related to the genus Bathymodiolus), tiny vesicomyid clams, and vestimentiferan clusters in which animals are up to 2m long. There is a high density of sponges (three species), bryozoans and gorgonians. Other taxa occasionally associated with this assemblage include galatheid crabs, anemones, holothurians and possibly pennatulaceans which are numerous at boundaries of the site. A typical deep-sea fauna of fish, asteroids and ophiuroids is also scattered about the vicinity. This seep community is distributed as a band of discontinuous colonies, 40–50m in width and more than 10km in length. Living colonies alternate with accumulations shells and bare areas. The substratum is hard mud covered by a ferrugineous crust and cemented by carbonates. The early diagenetic character of the bottom and the occurrence of biological assemblages indicate that seepages of sulphur and/or methane, which have been revealed along the decollement zone of the Barbados prism, must diffuse through the sea floor. These biological and geological features are located on the crests of NE-SW anticlinal ridges and mud volcanoes which have been built by diaprism.  相似文献   

13.
In and around the beds of vesicomyid clam (Calytogena soyoae) located off Hatsushima Island in Sagami Bay, central Japan, hydrogen sulfide concentration in bottom water and interstitial water was measured every 10 cm from just above seafloor to 40 cm deep usingin situ separative dialysis bags. While hydrogen sulfide over 0.01 mmol/kg was not measured from the seawater just above the dense clam beds, the concentration of hydrogen sulfide increased rapidly below 10 cm deep. The results indicate that the habit of the clam is correlated with high concentration of hydrogen sulfide contained in pore waters of sediments between depths of 10 and 20 cm from the bottom surface. Concentrations of hydrogen sulfide ranging from approximately 0.05 mmol/kg to 0.6 mmol/kg might be suitable requirement for the habitat ofC. soyoae.  相似文献   

14.
平端深海偏顶蛤(Gigantidasplatifrons)是广泛分布于西太平洋深海热液和冷泉生态系统中的优势种和共有种,也是深海化能生态系统中重要的生境营造种。以南海F冷泉平端深海偏顶蛤为研究对象,基于贝壳日生长轮方法,分析了平端深海偏顶蛤的年龄与生长速率,建立了其年龄与贝壳壳长关系的生长方程。同时,结合获得的生长方程,分析了南海F冷泉采样点(119.285 6°E,22.115 4°N)平端深海偏顶蛤种群的年龄结构。研究结果表明,采样获得的平端深海偏顶蛤种群的最大壳长为11.4 cm,最大寿命为13.5龄。研究区域的贝壳长度分布集中在4～7 cm,占比60%;年龄分布集中在2～4龄,占比49.7%。相关研究结果为进一步开展平端深海偏顶蛤的生长研究提供了基础数据,有助于深入了解冷泉区域的种群动态变化规律。  相似文献   

15.
Near the Azores Triple Junction as the Azores Plateau is approached, the ridge axis becomes shallower; its depth decreases from ca. 2400 m in the Rainbow vent field (36°13′N) to ca. 850 m in the Menez Gwen vent field (37°35′N). In this area, extensive mussel beds of the mytilid Bathymodiolus azoricus dominate the hydrothermal vent fauna, along with populations of three shrimps (Rimicaris exoculata, Mirocaris fortunata and Chorocaris chacei). The main physical and chemical characteristics of the vent habitat were studied by discrete sampling, in situ analysis and sediment trap moorings. The vent fauna is distributed along a variable band where the vent fluids and seawater mix, with R. exoculata living in the most concentrated areas and Bathymodiolus azoricus in the most diluted zones. Various non-endemic species live at the border of the vent field. The variations observed in structure and composition of the communities along the depth gradient are most likely due to changes in vent fluid toxicity (metallic and sulphide content) and suspended mineral particles, which render the fluids harsher for species living there. The main faunal differences observed between Lucky Strike and Menez Gwen hydrothermal fields are due to an impoverishment in the hydrothermal endemic species and to the penetration of bathyal species. The comparison of the three studied vent fields suggests the existence of a succession of several biogeographic islands rather than a single province.  相似文献   

16.
Vesicomyid clams, vestimentiferans, and some bathymodiolin mussels from hydrothermal vents and cold seeps possess thiotrophic endosymbionts, high levels of hypotaurine and, in tissues with symbionts, thiotaurine. The latter, a product of hypotaurine and sulfide, may store and/or transport sulfide non‐toxically, and the ratio to hypotaurine plus thiotaurine (Th/[H + Th]) may reflect an animal's sulfide exposure. To test this, we analyzed seep and vent animals with in situ sulfide measurements. Calyptogena kilmeri clams occur at high‐sulfide seeps in Monterey Canyon, while C. (Vesicomya) pacifica clams occur at seeps with lower levels but take up and metabolize sulfide more effectively. From one seep where they co‐occur, both had gill thiotaurine contents at 22–25 mmol kg−1 wet mass, and while C. (V.) pacifica had a higher blood sulfide level, it had a lower Th/[H + Th] (0.39) than C. kilmeri (0.63). However, these same species from different seeps with lower sulfide exposures had lower ratios. Bathymodiolus thermophilus [East Pacific Rise (EPR 9°50′ N)] from high‐(84 μm ) and a low‐(7 μm ) sulfide vents had gill ratios of 0.40 and 0.12, respectively. Trophosomes of Riftia pachyptila (EPR 9°50′ N) from medium‐(33 μm ) and low‐(4 μm ) sulfide vents had ratios of 0.23 and 0.20, respectively (not significantly different). Ridgeia piscesae vestimentiferans (Juan de Fuca Ridge) have very different phenotypes at high‐ and low‐sulfide sites, and their trophosomes had the greatest differences: 0.81 and 0.04 ratios from high‐ and low‐sulfide sites, respectively. Thus Th/[H + Th] may indicate sulfide exposure levels within species, but not in interspecies comparisons, possibly due to phylogenetic and metabolic differences. Total H + Th was constant within each species (except in R. piscesae); the sum may indicate the maximum potential sulfide load that a species faces.  相似文献   

17.
Although there is a growing body of evidence indicating benthic foraminifera inhabit hydrocarbon and cold seep environments, biochemical and ultrastructural data on seep foraminiferal communities are not available. Therefore, sediments collected from cold seeps in Monterey Bay, CA (900–1000 m), were examined for the presence of live benthic foraminifera. Results from three independent methods (ATP assay, ultrastructural analysis, rose Bengal staining) indicate that certain species inhabit the Clam Flat and Clam Field seeps. Abundances in our seep samples were lower than in comparable non-seep sites, although not atypical for these bathyal depths. Of 38 species represented at these two seep sites by cytoplasm-containing specimens, only Spiroplectammina biformis was restricted to the seep environment. However, because S. biformis is also known from non-seep sites in other areas, it should not be considered as endemic to seeps. Ultrastructural studies show abundant peroxisomes in seep specimens, which may allow inhabitation of such environments. One specimen of Uvigerina peregrina had prokaryotes nestled in test pores, suggesting that bacteria may play a role in the survival of foraminifera in this seep environment.  相似文献   

18.
Quantitative collections of tubeworm- and mussel-associated communities were obtained from 3 cold seep sites in the deep Gulf of Mexico: in Atwater Valley at 1890 m depth, in Alaminos Canyon at 2200 m depth, and from the Florida Escarpment at 3300 m depth. A total of 50 taxa of macro- and megafauna were collected including 2 species of siboglinid tubeworms and 3 species of bathymodiolin mussels. In general, the highest degree of similarity was between communities collected from the same site. Most of the dominant families at the well-characterized upper Louisiana slope seep sites of the Gulf of Mexico were present at the deep sites as well; however, there was little overlap at the species level between the upper and lower slope communities. One major difference in community structure between the upper and lower slope seeps was the dominance of the ophiuroid Ophioctenella acies in the deeper communities. The transition between upper and lower slope communities appears to occur between 1300 and 1700 m based on the number of shared species with the Barbados seeps at either end of this depth range. Seep communities of the deep Gulf of Mexico were more similar to the Barbados Accretionary Prism seep communities than they were to either the upper slope Gulf of Mexico or Blake Ridge communities based on numbers of shared species and Bray–Curtis similarity values among sites. The presence of shared species among these sites suggests that there is ongoing or recent exchange among these areas. An analysis of bathymodioline mussel phylogeography that includes new collections from the west coast of Africa is presented. This analysis also suggests recent exchange across the Atlantic equatorial belt from the Gulf of Mexico to the seeps of the West Nigerian margin.  相似文献   

19.
Associations between scale‐worms and giant mussels are common constituents of hydrothermal vent and cold seep ecosystems, but very little is known about their nature and ecology. Here, we analyze the ecological characteristics of the associations between Branchipolynoe seepensis, an obligate symbiotic polychaete, and their host mytilid mussels Bathymodiolus puteoserpentis and B. azoricus inhabiting hydrothermal vent fields on the Mid‐Atlantic Ridge. Infested mussels generally harbored a single symbiont (<4% had two to six worms). Infestation rate varied from 7.2% to 76.5%, increasing with mussel size, and was significantly lower for B. puteoserpentis. Symbiont density at Lucky Strike ranged between 1071 and 1191 individuals m−2. Female symbiont size was always positively correlated with host size, while only males and juveniles from small mussels showed the same trend. This suggested a relatively long‐lasting host/symbiont association for females and short‐lasting association with successive reproductive migrations for adult males. The sex ratio of symbionts was always biased in favor of females. Males were smaller and more slender than females and had one mode in their size distributions, whereas females typically had three or more modes, suggestive of a longer life span in females. Between 59.1% and 72.2% of mussels had damaged soft tissues with substantially higher incidence of trauma in infested ones, suggesting that symbionts may cause trauma. The symbionts also induce tunnel‐like structures among the ctenidia, indicating fidelity to a particular location inside the host. Based on our data, together with the fact that infested mussels became relatively wider than non‐infested ones, this association is considered parasitic (likely kleptoparasitic). Our data, together with those from previous studies, allowed us to define the main life‐history traits of B. seepensis: (i) the relationship with their host is parasitic, (ii) the association begins at the smallest mytilid size classes, (iii) there is sexual dimorphism in body size, (iv) sex ratio deviates from 1:1 in favor of females, (v) fertilization occurs through temporal pairing and pseudocopulation, (vi) sperm are stored by females, (vii) eggs are large (likely lecithotrophic or with direct development), (viii) females have a longer life span than males, (ix) adult males may be semalparous, undertaking reproductive migrations followed by a short period of pairing and then death, and (x) females have a semi‐continuous iteroparous reproductive cycle.  相似文献   

20.
细菌通过趋化系统感知和响应外界环境变化并进行趋利避害, 因此很多细菌进化出了非常复杂多样的趋化系统以适应不同的生态位。Epsilon-变形菌广泛存在于自然界中, 能适应不同的生态环境, 特别是深海Epsilon-变形菌能适应深海热液口和冷泉等极端环境, 其趋化系统可能有其独特之处。通过BlastP和MIST数据库分析, 我们发现大部分深海Epsilon-变形菌拥有F3类型趋化系统且具有单拷贝的双功能域融合蛋白CheV。此外, 一种特殊结构域CZB (C-terminal Zinc-Binding) 存在于深海Epsilon-变形菌的趋化受体蛋白中, 而且在部分深海Epsilon-变形菌的趋化受体中还发现并鉴定了一种新的结构域CZB-like结构域。文章以Epsilon-变形菌的模式菌株空肠弯曲杆菌(Campylobacter jejuni 81-176)为研究对象, 用细菌双杂交试验证实了CheV能与所有含MA结构域的趋化受体相互作用。用电感耦合等离子体质谱(ICP-Mass)技术证实了CZB-like结构域不能结合Zn离子, 但细菌双杂交试验证实它能介导趋化受体Tlp9与CheV的互作。  相似文献   

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