首页 | 本学科首页   官方微博 | 高级检索  
相似文献
 共查询到20条相似文献，搜索用时 31 毫秒
1.
New large-scale laboratory data are presented on the influence of long waves, bichromatic wave groups and random waves on sediment transport in the surf and swash zones. Physical model testing was performed in the large-scale CIEM wave flume at UPC, Barcelona, as part of the SUSCO (swash zone response under grouping storm conditions) experiment in the Hydralab III program (Vicinanza et al., 2010). Fourteen different wave conditions were used, encompassing monochromatic waves, bichromatic wave groups and random waves. The experiments were designed specifically to compare variations in beach profile evolution between monochromatic waves and unsteady waves with the same mean energy flux. Each test commenced with approximately the same initial profile. The monochromatic conditions were perturbed with free long waves, and then subsequently substituted with bichromatic wave groups with different bandwidth and with random waves with varying groupiness. Beach profile measurements were made at half-hourly and hourly intervals, from which net cross-shore transport rates were calculated for the different wave conditions. Pairs of experiments with slightly different bandwidth or wave grouping show very similar net cross-shore sediment transport patterns, giving high confidence to the data set. Consistent with recent small-scale experiments, the data clearly show that in comparison to monochromatic conditions the bichromatic wave groups reduce onshore transport during accretive conditions and increase offshore transport during erosive conditions. The random waves have a similar influence to the bichromatic wave groups, promoting offshore transport, in comparison to the monochromatic conditions. The data also indicate that the free long waves promote onshore transport, but the conclusions are more tentative as a result of a few errors in the test schedule and modifications to the setup which reduced testing time. The experiments suggest that the inclusion of long wave and wave group sediment transport is important for improved near-shore morphological modeling of cross-shore beach profile evolution, and they provide a very comprehensive and controlled series of tests for evaluating numerical models. It is suggested that the large change in the beach response between monochromatic conditions and wave group conditions is a result of the increased significant and maximum wave heights in the wave groups, as much as the presence of the forced and free long waves induced by the groupiness. The equilibrium state model concept can provide a heuristic explanation of the influence of the wave groups on the bulk beach profile response if their effective relative fall velocity is larger than that of monochromatic waves with the same incident energy flux.  相似文献   

2.
《Coastal Engineering》2005,52(1):1-23
We develop solutions for the transport of suspended sediment by a single swash event following the collapse of a bore on a plane beach, and we investigate the morphodynamical role that such transport may play. Although the intrinsic asymmetry between uprush and backwash velocities tends to encourage the export of sediment, we find that swash events may be effective in distributing across the swash zone much or all of the sediment mobilised by bore collapse; additionally, settling lag effects may promote a weak onshore movement of sediment. We quantify both effects in terms of the properties of the sediment and of the swash event, and comment on the relationship between our findings and recent field studies of swash zone sediment transport.  相似文献   

3.
A critical review of conceptual and mathematical models developed in recent decades on sediment transport in the swash zone is presented. Numerous studies of the hydrodynamics and sediment transport in the swash zone in recent years have pointed out the importance of swash processes in terms of science advancement and practical applications. Evidently, the hydrodynamics of the swash zone are complex and not fully understood. Key hydrodynamic processes include both high-frequency bores and low-frequency infragravity motions, and are affected by wave breaking and turbulence, shear stresses and bottom friction. The prediction of sediment transport that results from these complex and interacting processes is a challenging task. Besides, sediment transport in this oscillatory environment is affected by high-order processes such as the beach groundwater flow. Most relationships between sediment transport and flow characteristics are empirical, based on laboratory experiments and/or field measurements. Analytical solutions incorporating key factors such as sediment characteristics and concentration, waves and coastal aquifer interactions are unavailable. Therefore, numerical models for wave and sediment transport are widely used by coastal engineers. This review covers mechanisms of sediment transport, important forcing factors, governing equations of wave-induced flow, groundwater interactions, empirical and numerical relations of cross-shore and longshore sediment transport in the swash zone. Major advantages and shortcomings of various numerical models and approaches are highlighted and reviewed. These will provide coastal modelers an impetus for further detailed investigations of fluid and sediment transport in the swash zone.  相似文献   

4.
The morphological changes of multiple intertidal bars (ridges) on a macrotidal beach were examined under low-energy wave conditions during a spring-to-spring tidal cycle. The morphological response was coupled to the tidal water level variations and related residence times for swash processes and surf (breaking waves and bores) over the cross-shore profile. Spring tides induced a large spatial variation in water lines and small residence times for distinct processes. Neap tides narrowed the intertidal area and increased the time for certain processes to work on the sediment at one location. The observed morphological changes could be coupled to the stagnation of processes at a certain bar crest position. The action of surf (breaking waves and bores) played the major role in the onshore migration of the intertidal bars and the simultaneous erosion of the seaward flank. Swash action, responsible for the generation and migration of intertidal bars in microtidal settings, was not the dominant process in causing the observed morphological changes. Intertidal ridges on macrotidal beaches cannot be considered swash bars as suggested by most previous investigations into these morphological features.  相似文献   

5.
Hydrodynamics and sediment transport in the nearshore zone were modeled numerically taking into account turbulent unsteady flow. The flow field was computed using the Reynolds Averaged Navier–Stokes equations with a k–ε turbulence closure model, while the free surface was tracked using the Volume-Of-Fluid technique. This hydrodynamical model was supplemented with a cross-shore sediment transport formula to calculate profile changes and sediment transport in the surf and swash zones. Based on the numerical solutions, flow characteristics and the effects of breaking waves on sediment transport were studied. The main characteristic of breaking waves, i.e. the instantaneous sediment transport rate, was investigated numerically, as was the spatial distribution of time-averaged sediment transport rates for different grain sizes. The analysis included an evaluation of different values of the wave friction factor and an empirical constant characterizing the uprush and backwash. It was found that the uprush induces a larger instantaneous transport rate than the backwash, indicating that the uprush is more important for sediment transport than the backwash. The results of the present model are in reasonable agreement with other numerical and physical models of nearshore hydrodynamics. The model was found to predict well cross-shore sediment transport and thus it provides a tool for predicting beach morphology change.  相似文献   

6.
《Marine Geology》2004,203(1-2):109-118
Spatial variations in sediment load in the swash uprush and textural properties of sediment in transport were evaluated to investigate the mechanisms responsible for sediment transport during wave uprush. Four streamer traps were deployed at 2.0-m intervals across the swash zone of a sheltered, microtidal sandy beach at Port Beach, Western Australia, over a 4-day period. During these trapping experiments, offshore significant wave heights were 0.3–0.5 m and wave periods were about 10 s. The average width of the uprush zone was 6.9 m and the average uprush duration was 5.9 s. Cross-shore distributions of sediment load for 70 uprush events reveal a maximum in sediment load landward of the base of the swash (at about 20% of swash width) during single events and a maximum closer to mid-swash (at about 40% of swash width) during multiple events characterized by swash interactions. Settling velocity distributions of trap samples during individual uprush events are similar to distributions found on the beach surface, with the lowest settling velocities (finest sediments) near the base of the swash zone and maximum settling velocities (coarsest sediments) around the mid-swash position. It was found that sediment transport during wave uprush occurs through two distinct mechanisms: (1) sediment entrainment during bore collapse seaward of the base of the swash zone and subsequent advection of this bore-entrained sediment up the beach by wave uprush; and (2) in situ sediment entrainment and transport induced by local shear stresses during wave uprush. Both mechanisms are considered important, but the first mechanism is considered most significant during the early stages of wave uprush when sediment is transported mainly in suspension, while the second mechanism is likely to dominate the mid- to later stages of wave uprush when sediment is transported mainly by sheet flow. The relative importance of the two mechanisms will vary between different beaches with the morphodynamic state of the beach (reflective versus dissipative) expected to play a major role.  相似文献   

7.
Time-series of nearbed horizontal flow velocities and suspended sediment concentrations obtained from a colocated electromagnetic current meter (EMCM) and optical backscatter sensor (OBS), respectively, are used to examine the relative importance of steady and fluctuating components to the total sediment transport over a full tidal cycle on a macrotidal, intermediate beach (Spurn Head, UK). Fluctuating sediment fluxes are decomposed into gravity and infragravity contributions using co-spectral techniques. The relative importance of the oscillatory (gravity and infragravity) and steady (mean) transport components to the total sediment transport is analysed throughout the tidal cycle.

A continuum of 34 discrete suspended sediment-cross-shore velocity co-spectra are computed over a full tidal cycle for the OBS and EMCM measurements 0.10 m above the bed. These net transport spectra vary greatly both with cross-shore location and tidal state. In particular, a marked asymmetry in transport processes is evident between the flood and ebb tides, with high levels of sediment resuspension and transport occurring on the ebbing tide approximately two hours after high water (just seaward of the breakpoint). At this time the dominant transport was directed offshore (co-spectral peak, 0.04 kg/m2/s) at incident wave frequency.

Typical patterns are observed in transport spectra outside the surf zone and within the inner surf zone. Outside the narrow surf zone cross-shore transport spectra show weak offshore transport (co-spectral peak = 0.002 kg/m2/s) associated with bound long waves and stronger onshore transport (co-spectral peak = 0.006 kg/m2/s) at incident wave frequencies. Conversely, co-spectra computed within the inner surf zone show the offshore sediment fluxes (spectral peak = 0.010 kg/m2/s) at infragravity frequencies to be greater in magnitude than the corresponding onshore transport (co-spectral peak = 0.008 kg/m2/s) occurring at incident wave frequencies.  相似文献   


8.
《Coastal Engineering》2006,53(4):335-347
This paper investigates cross-shore profile changes of gravel beaches, with particular regard to discussing the tendency for onshore transport and profile steepening in the swash zone. The discussion includes observed morphological changes on a gravel beach from experimental investigations at the Large Wave Flume (GWK) in Hanover, Germany. During the tests all the profile changes occurred in the swash zone, resulting in erosion below the still water line (SWL) and formation of a berm above the SWL. We investigate the profile evolution evaluating the transport rates from a bed load sediment transport formulation coupled with velocities calculated from a set of Boussinesq equations that have been validated for its use in the surf and swash zones [Lynett, P.J., Wu, T.-R., and Liu, L.-F., P., 2002. Modelling wave runup with depth-integrated equations. Coastal Engineering, 46, 89–107; Otta, A.K., and Pedrozo-Acuña, A., 2004. Swash boundary and cross-shore variation of horizontal velocity on a slope. In: J.M. Smith (Editor), Proceedings 29th International Conference on Coastal Engineering. World Scientific, Lisbon, Portugal, pp. 1616–1628]. We discuss the influence of bottom friction on the predicted profiles, using reported friction factors from experimental studies. It is shown that the use of a different friction factor within a realistic range in each phase of the swash (uprush and backwash) improves prediction of the beach profiles, although quantitative agreement between the measured and computed profile evolutions is not satisfactory. Furthermore, if the friction factor and the transport efficiency (C) of the sediment transport formulation are kept the same in the uprush and backwash, accurate representation of profile evolution is not possible. Indeed, the features of the predicted profiles are reversed. However, when the C parameter is set larger during the uprush than during the backwash, the predicted profiles are closer to the observations. Differences between the predicted profiles from setting non-identical C-values and friction factors for the swash phase, are believed to be linked to both the infiltration effects on the flow above the beachface and the more accelerated flow in the uprush.  相似文献   

9.
Novel laboratory experiments and numerical modelling have been performed to study the advection scales of suspended sediment in the swash zone. An experiment was designed specifically to measure only the sediment picked up seaward of the swash zone and during bore collapse. The advection scales and settling of this sediment were measured during the uprush along a rigid sediment-free beach face by a sediment trap located at varying cross-shore positions. Measurements were made using a number of repeated solitary broken waves or bores. Approximately 25% of the pre-suspended sediment picked up by the bores reaches the mid-swash zone (50% of the horizontal run-up distance), indicating the importance of the sediment advection in the lower swash zone. The pre-suspended sediment is sourced from a region seaward of the shoreline (still water line) which has a width of about 20% of the run-up distance. An Eulerian–Lagrangian numerical model is used to model the advection scales of the suspended sediment. The model resolves the hydrodynamics by solving the non-linear shallow water equations in an Eulerian framework and then solves the advection–diffusion equation for turbulence and suspended sediment in a Lagrangian framework. The model provides good estimates of the measured mass and distribution of sediment advected up the beach face. The results suggest that the correct modelling of turbulence generation prior to and during bore collapse and the advection of the turbulent kinetic energy into the lower swash is important in resolving the contribution of pre-suspended sediment to the net sediment transport in the swash zone.  相似文献   

10.
The morphology, bedforms and hydrodynamics of Merlimont beach, in northern France, characterised by intertidal bars and a spring tidal range of 8.3 m, were surveyed over a 10-day experiment with variable wave conditions that included a 2-day storm with significant wave heights of up to 2.8 m. The beach exhibited two pronounced bar-trough systems located between the mean sea level and low neap tide level. Waves showed a cross-shore depth modulation, attaining maximum heights at high tide. The mean current was characterised dominantly by strong tide-induced longshore flows significantly reinforced by wind forcing during the storm, and by weaker, dominantly offshore, wave-induced flows. Vertical tidal water-level variations (tidal excursion rates) showed a bimodal distribution with a peak towards the mid-tide position and low rates near low and high water. The two bar-trough systems in the mid-tide zone remained stable in position during the experiment but showed significant local change. The absence of bar migration in spite of the relatively energetic context of this beach reflects high macro-scale bar morphological lag due to a combination of the large vertical tidal excursion rates in the mid-tide zone, the cross-shore wave structure, and the pronounced dual bar-trough system. The profile exhibited a highly variable pattern of local morphological change that showed poor correlation with wave energy levels and tidal excursion rates. Profile change reflected marked local morphodynamic feedback effects due mainly to breaks in slope associated with the bar-trough topography and with trough activity. Change was as important during low wave-energy conditions as during the storm. Strong flows in the entrenched troughs hindered cross-shore bar mobility while inducing longshore migration of medium-sized bedforms that contributed in generating short-term profile change. The large size and location of the two pronounced bars in the mid-tide zone of the beach are tentatively attributed respectively to the relatively high wave-energy levels affecting Merlimont beach, and to the cross-shore increase in wave height hinged on tidal modulation of water depths. These two large quasi-permanent bars probably originated as essentially breakpoint bars and are different from a small bar formed by swash and surf processes in the course of the experiment at the mean high water neap tide level, which is characterised by a certain degree of tidal stationarity and larger high-tide waves.  相似文献   

11.
《Coastal Engineering》2005,52(7):633-645
New experimental laboratory data are presented on swash overtopping and sediment overwash on a truncated beach, approximating the conditions at the crest of a beach berm or inter-tidal ridge-runnel. The experiments provide a measure of the uprush sediment transport rate in the swash zone that is unaffected by the difficulties inherent in deploying instrumentation or sediment trapping techniques at laboratory scale. Overtopping flow volumes are compared with an analytical solution for swash flows as well as a simple numerical model, both of which are restricted to individual swash events. The analytical solution underestimates the overtopping volume by an order of magnitude while the model provides good overall agreement with the data and the reason for this difference is discussed. Modelled flow velocities are input to simple sediment transport formulae appropriate to the swash zone in order to predict the overwash sediment transport rates. Calculations performed with traditional expressions for the wave friction factor tend to underestimate the measured transport. Additional sediment transport calculations using standard total load equations are used to derive an optimum constant wave friction factor of fw = 0.024. This is in good agreement with a broad range of published field and laboratory data. However, the influence of long waves and irregular wave run-up on the overtopping and overwash remains to be assessed. The good agreement between modelled and measured sediment transport rates suggests that the model provides accurate predictions of the uprush sediment transport rates in the swash zone, which has application in predicting the growth and height of beach berms.  相似文献   

12.
响应季节性波候作用的泥沙输运特征是研究弧形海滩地貌变化及港工建筑的重要内容。基于南湾弧形海滩实际测量的冬、夏各11条剖面高程变化资料,将其划分为低潮间带、低中潮带、中潮带、高潮间带、低冲流带、中冲流带及其海滩后滨等7带,在此基础上利用经验正交函数(EOF)方法对各个带的体积变化进行分析,结果表明:1)南湾弧形海滩的泥沙以单向输运为主,并具有季节性变化特征,其中冬季泥沙在东南浪作用下,自陆向海输运,夏季泥沙在西南浪作用下自海向陆输运;2)南湾弧形海滩的泥沙分别在高潮带与中潮带、低冲流带与中冲流带之间存在频繁的双向输运;3)南湾弧形海滩不同岸段泥沙的横向输运因岬角的遮蔽能力、地形以及波浪作用的方向而有所差异。  相似文献   

13.
The formation of beach megacusps along the shoreline of southern Monterey Bay, CA, is investigated using time-averaged video and simulated with XBeach, a recently developed coastal sediment transport model. Investigations focus on the hydrodynamic role played by the bay's ever-present rip channels. A review of four years of video and wave data from Sand City, CA, indicates that megacusps most often form shoreward of rip channels under larger waves (significant wave height (Hs) = 1.5–2.0 m). However, they also occasionally appear shoreward of shoals when waves are smaller (Hs ~ 1 m) and the mean water level is higher on the beach. After calibration to the Sand City site, XBeach is shown to hindcast measured shoreline change moderately well (skill = 0.41) but to overpredict the erosion of the swash region and beach face. Simulations with small to moderate waves (Hs = 0.5–1.2 m) suggest, similar to field data, that megacusps will form shoreward of either rip channels or shoals, depending on mean daily water level and pre-existing beach shape. A frequency-based analysis of sediment transport forcing is performed, decomposing transport processes to the mean, infragravity, and very-low-frequency (VLF) contributions for two highlighted cases. Results indicate that the mean flow plays the dominant role in both types of megacusp formation, but that VLF oscillations in sediment concentration and advective flow are also significant.  相似文献   

14.
Coastal groundwater systems can have a considerable impact on sediment transport and foreshore evolution in the surf and swash zones. Process-based modeling of wave motion on a permeable beach taking into account wave-aquifer interactions was conducted to investigate the effects of the unconfined coastal aquifer on beach profile evolution, and wave shoaling on the water table. The simulation first dealt with wave breaking and wave runup/rundown in the surf and swash zones. Nearshore hydrodynamics and wave propagation in the cross-shore direction were simulated by solving numerically the two-dimensional Navier–Stokes equations with a k–ε turbulence closure model and the Volume-Of-Fluid technique. The hydrodynamic model was coupled to a groundwater flow model based on SEAWAT-2000, the latter describing groundwater flow in the unconfined coastal aquifer. The combined model enables the simulation of wave-induced water table fluctuations and the effects of infiltration/exfiltration on nearshore sediment transport. Numerical results of the coupled ocean/aquifer simulations were found to compare well with experimental measurements. Wave breaking and infiltration/exfiltration increase the hydraulic gradient across the beachface and enhance groundwater circulation inside the porous medium. The large hydraulic head gradient in the surf zone leads to infiltration across the beachface before the breaking point, with exfiltration taking place below the breaking point. In the swash zone, infiltration occurs at the upper part of the beach and exfiltration at the lower part. The simulations confirm that beaches with a low water table tend to be accreted while those with a high water table tend to be eroded.  相似文献   

15.
Field experiments were conducted on a low-gradient, high-energy sandy beach (Truc Vert, France) and a steep, low-energy gravel beach (Slapton, UK) to examine alongshore-directed currents within the swash zone. At Truc Vert, data were collected over 33 tidal cycles with offshore significant wave heights of 1–4 m and periods of 5–12 s. At Slapton data were collected during 12 tides with wave heights of 0.3–1 m and periods of 4–9 s. The swash motion was predominantly at infragravity frequencies at Truc Vert and incident frequencies at Slapton.  相似文献   

16.
《Coastal Engineering》2001,43(1):25-40
Video-based swash motions from three studies (on two separate beaches) were analyzed with respect to theoretical swash trajectories assuming plane beach ballistic motions under quadratic friction. Friction coefficient values for both the uprush and backwash were estimated by comparing measured swash space–time trajectories to these theoretical expectations given an initial velocity and beach slope. Observations were made spanning high tides, and in one case, during a light rain. Analysis of over 4500 individual swash events showed that the uprush friction coefficient was nearly constant during all three studies with a mean value of roughly 0.007 and showed no trends over a tidal cycle. In contrast, backwash friction coefficient values varied over the tidal cycles ranging between 0.01 and 0.07 with minimum values corresponding to the highest tides. Although these values are close to the theoretical estimates based on a Law of the Wall formulation and values commonly referenced in the literature, these observations show a consistent tendency for backwash friction estimates to greatly exceed uprush friction estimates. The disparity between uprush and backwash friction coefficients can be partially attributed to the exclusion of a pressure gradient term in the ballistic model. However, results indicate that backwash friction coefficients adjusted to account for this effect may be three times larger than the uprush friction values during lower tides. This tidal dependence for backwash friction coefficients is attributed to a complex interaction between swash infiltration and entrained sediment loads. These findings imply that friction estimates (necessary for sediment transport calculations and hydrodynamic predictions) based solely on grain roughness may not be correct for backwash flows.  相似文献   

17.
The south-western shoreline along the entrance channel inside the Port of Richards Bay has experienced continued erosion. Four groynes were constructed to stabilise the shoreline. Monitoring of shoreline evolution provided valuable data on the accretion adjacent to two of the groynes and on the sediment transport rates at these groynes. Tides, beach slopes, winds, wave climate, current regime, and sand grain sizes were documented. The one site is “moderately protected” from wave action while the other is “protected” according to the Wiegel [Wiegel, R. L. (1964). Oceanographical engineering. Prentice Hall, Inc., Englewood Cliffs, NJ.] classification. The shoreline accreted progressively at the two groynes at 0.065 m/day and 0.021 m/day respectively. The shorelines accreted right up to the most seaward extremity of the groynes. Equilibrium shorelines were reached within about 3.5 years to 4 years, which compare well with other sites around the world. The mean wave incidence angle is large and was found to be about 22°. The median sand grain sizes were 0.33 mm and 0.37 mm. The groynes acted as total traps, the beach surveys were extended to an adequate depth, and cross-shore sediment transport did not cause appreciable net sand losses into the entrance channel. The net longshore transport rate along the study area, which is north-westbound, is only slightly lower than the gross longshore transport. The actual net longshore transport rates are 18 000 m3/year and 4 600 m3/year respectively at the two groynes. A rocky area limits the availability of sand at one groyne. There is fair agreement between the predicted and measured longshore transport rates at the other groyne.  相似文献   

18.
  相似文献   

19.
Three sediment transport studies using tracers were performed at Ancão Inlet (southern Portugal). The objectives of the experiments were to understand the sediment transport pathways and to determine their magnitudes on the updrift margin of an inlet. In order to apply the traditionally used Space Integration Methodology to the tracer experiments, adaptations were required. The study area was divided into four morphologically defined sectors and this was found to be a key factor for the applicability of tracers in a complex area. The four sectors are as follows: sector A is the straight part of the updrift beach; sector B is the upper area of the swash platform; sector C is the lower area of the swash platform; and sector D represents the inner parts of the inlet margin. The integrated analysis of all collected data (forcing mechanisms, tracer distribution and topographic evolution) led to the determination of the sediment pathways. A semi-quantitative conceptual model was developed in order to explain the sediment transport pathways and magnitudes that a known mass of sediment would follow after arriving at the swash platform. It was found that the areas with the largest sediment accumulation were sectors B and C, while almost no sediment was retained in sector D, which experienced significant erosion. According to the model, 53% of the initial mass of sediments remain in the system after two tidal cycles. It is hypothesised that sediment losses are caused by sediment transport towards the ebb delta and by sediment bypassing occurring from the ebb delta to the downdrift beach through swash bar processes. The herein defined conceptual model represents a useful tool that could be applied to other tidal inlets under similar conditions, facilitating sediment budget studies around tidal inlets.  相似文献   

20.
Many macrofaunal species inhabiting exposed sandy beaches are dependent on the swash for their nutrition and migration and are highly adapted to the harsh physical conditions of the swash. The most important physical factors that determine the distribution and behaviour of swash related fauna, next to the swash itself, are sediment grain size and beach slope. Crucial swash parameters are swash period and swash velocity. Studying the influence of these factors on the animals in the field is often very difficult and it is almost impossible to identify which factor causes what effect. Crucial knowledge about the direct role of the swash itself is lacking. Therefore, a device that generates swash waves on an artificial beach under laboratory conditions was designed: the swash rig. In the swash rig, full and independent control on sediment grain size, beach slope, swash velocity and swash period is present. This will allow us to do a variety of experiments on the influence of each of these factors, independently or combined, on swash fauna. In one such experiment, Olivella semistriata – a dominant surfing gastropod on Ecuadorian sandy beaches – was placed in the swash rig during rising and falling tide and subjected – under constant conditions – to an equal wave regime. During falling tide, and in absence of any tidal cue, almost all specimens moved downshore, as they would in the field; hardly any specimens moved upshore. During upcoming tide, however, there was noticeable upward migration in the swash rig, and half of the runs showed a net upward migration. Contrary to the common understanding that the behaviour of sandy beach molluscs is entirely environmentally driven, this experiment hints at the presence of an endogenous circatidal clock, which is used to direct the tidal migration of the species.  相似文献   

设为首页 | 免责声明 | 关于勤云 | 加入收藏

Copyright©北京勤云科技发展有限公司  京ICP备09084417号